02439nas a2200133 4500008004100000022004600041245004300087210003900130260006400169300001200233520200200245100001902247856003902266 2019 eng d a978-952-11-4973-3, ISBN 978-952-11-4974-000aTuhatjalkaiset - Myriapods - Myriapoda0 aTuhatjalkaiset Myriapods Myriapoda aHelsinkibYmpäristöministeriö & Suomen ympäristökeskus a340-3433 a

Viittausohje, kun viitataan yksittäiseen Punaisen listan lukuun: Vahtera, V. 2019. Tuhatjalkaiset. Julk.: Hyvärinen, E., Juslén, A., Kemppainen, E., Uddström, A. & Liukko, U.-M. (toim.) 2019. Suomen lajien uhanalaisuus – Punainen kirja 2019. Ympäristöministeriö & Suomen ympäristökeskus. Helsinki. S. 340-343. http://hdl.handle.net/10138/299501

Reference of a separate chapter of the Red List: Vahtera, V. 2019. Myriapods. In: Hyvärinen, E., Juslén, A., Kemppainen, E., Uddström, A. & Liukko, U.-M. (eds.) 2019. The 2019 Red List of Finnish Species. Ministry of the Environment & Finnish Environment Institute. Helsinki. P. 340-343. http://hdl.handle.net/10138/299501

Summary: Myriapoda are a diverse group of terrestrial arthropods, some of which are herbivores, some predators and some decomposers. There are four classes of Myriapoda: centipedes (Chilopoda), millipedes (Diplopoda), pauropods (Pauropoda), and symphylans (Symphyla). In Finland, a total of 63 species have been found: 21 centipede, 28 millipede, eight pauropod, and six symphylan species. In total, there are ten red-listed species: the critically endangered (CR) Geophilus carpophagus, which was classified as Regionally Extinct (RE) in the two previous assessments, and the Near Threatened (NT) species G. electricus, G. truncorum and Julus scanicus. These species only have one known locality each, and no new observations have been made of the three Near Threatened (NT) species for a long time. Six species have been categorised as Data Deficient (DD). Many species were classified as Not Evaluated (NE) due to insufficient information on their occurrence. While no monitoring of Myriapoda has been organised in Finland, the information on the spread of the species in this country could be updated by organising a monitoring study of a few years in duration.

1 aVahtera, Varpu uhttp://hdl.handle.net/10138/29950108893nam a2200337 4500008004100000022004600041245005500087210005100142260006400193300000800257520789800265653001508163653002808178653001708206653000908223653002108232653001908253653001308272653001408285653001908299653002208318653002308340653002208363653002608385100002108411700001808432700002108450700002208471700002308493856003908516 2019 eng d a978-952-11-4973-3, ISBN 978-952-11-4974-000aSuomen lajien uhanalaisuus – Punainen kirja 20190 aSuomen lajien uhanalaisuus Punainen kirja 2019 aHelsinkibYmpäristöministeriö & Suomen ympäristökeskus a7043 a

Tiivistelmä: Julkaisussa esitellään Suomen lajien viides uhanalaisuusarviointi. Suomen lajimäärän arvioidaan nykyään olevan vähintään 48 000. Riittävät tiedot uhanalaisuusarviointiin oli 22 418 lajista tai alemmasta taksonista, mikä on noin 47 % lajistosta. Uhanalaisiksi arvioitiin 2 668 lajia tai alempaa taksonia eli noin 11,9 % arvioiduista lajeista. Punaisen listan lajeja, joihin kuuluvat uhanalaisten lisäksi hävinneet, silmälläpidettävät ja puutteellisesti tunnetut on yhteensä 6 683 (29,8 %). Arvioiduista lajeista äärimmäisen uhanalaisia (CR) on 489, erittäin uhanalaisia (EN) 918, vaarantuneita (VU) 1 260, silmälläpidettäviä (NT) 1 912, puutteellisesti tunnettuja (DD) 1 792, hävinneitä (RE) 312 ja elinvoimaisia (LC) 15 735. Arviointi tehtiin Kansainvälisen luonnonsuojeluliiton (IUCN) ohjeiden ja kriteerien mukaan. IUCN:n alueellisen uhanalaisuusarvioinnin kriteerit perustuvat lajien havaittuun, arvioituun, pääteltyyn tai ennustettuun populaatioiden pienenemiseen, levinneisyys- tai esiintymisalueen kokoon, pieneen ja jatkuvasti taantuvaan tai hyvin pieneen populaatioon tai kvantitatiiviseen analyysiin. Lisäksi Suomessa on dokumentoitu lajien elinympäristöt sekä uhanalaisuuden syyt ja uhkatekijät. Enemmistö uhanalaisista lajeista elää metsissä (31 %) sekä perinneympäristöissä ja muissa ihmisen muuttamissa ympäristöissä (24 %). Uhanalaisten osuus kunkin elinympäristön arvioiduista lajeista on korkein tunturipaljakalla (37,9 %). Uhanalaisten lajien määrä on suurin hemiboreaalisella ja eteläboreaalisella vyöhykkeellä Etelä-Suomessa. Julkaisussa vertaillaan arvioinnin tuloksia vuoden 2010 tuloksiin esimerkiksi uhanalaisuusindeksin avulla. Lisäksi julkaisussa tarkastellaan vuoden 2010 uhanalaisuusarvioinnin ohjausryhmän ehdotusten toteutumista. Arvioinnin tulosten pohjalta Eliölajien uhanalaisuuden arvioinnin ohjausryhmä (LAUHA) esittää 13 toimenpide-ehdotusta uhanalaisten lajien suojelun, hoidon, tutkimuksen ja seurannan järjestämiseksi ja rahoittamiseksi. Arvioinnit on tehty eliöryhmäkohtaisissa työryhmissä ja työn tulokset esitellään eliöryhmittäin 39 luvussa.

Viittausohje, kun viitataan koko julkaisuun: Hyvärinen, E., Juslén, A., Kemppainen, E., Uddström, A. & Liukko, U.-M. (toim.) 2019. Suomen lajien uhanalaisuus – Punainen kirja 2019. Ympäristöministeriö & Suomen ympäristökeskus. Helsinki. 704 s. http://hdl.handle.net/10138/299501

Sammandrag: I denna bok presenteras resultat av Finlands femte hotbedömning. Finlands artantal uppskattas för tillfället till minst 48 000. Tillräckliga uppgifter för hotbedömning fanns om 22 418 arter eller lägre taxa, vilket utgör 47 % av alla arter. Av dessa bedömdes 2 667 arter eller lägre taxa vara hotade, d.v.s. 11,9 %. Rödlistade arter, som förutom hotade arter även innefattar försvunna arter, nära hotade arter och arter som det råder kunskapsbrist om, uppgår till 6 683 (29,8 %). Av de bedömda arterna är 498 akut hotade (CR), 918 starkt hotade (EN), 1 260 sårbara (VU), 1 912 nära hotade (NT), 1 792 med kunskapsbrist (DD), 312 försvunna (RE) och 15 735 livskraftiga. Hoten har bedömts på basis av Internationella Naturvårdsunionens (IUCN) regionala hotbedömningskriterier. Kriterierna grundar sig på observerad, deduktiv eller estimerad minskning av populationer eller utbrednings- och förekomstområden, en liten och kontinuerligt minskande eller en mycket liten population eller på kvantitativ analys. Därutöver har arternas livsmiljö, hotorsak och hotfaktorer i Finland dokumenterats. Majoriteten av de hotade arterna lever i skogar (31 %), vårdbiotoper och andra kulturmiljöer (24 %). Antalet hotade arter är högst inom den hemiboreala och sydboreala zonen i södra Finland. Andelen hotade arter i en specifik habitattyp är högst i kalfjället(37,9 %). I publikationen jämförs resultaten av hotbedömningen med resultaten från år 2010 t.ex. med hjälp av rödlisteindexet. Vid sidan av bedömningsresultaten beskrivs de mest påtagliga administrativa och juridiska åtgärder som vidtagits för artskydd samt forskning och övervakning sedan den föregående bedömningen. Hotbedömningens styrgrupp (LAUHA ) föreslår 13 åtgärder för organisering och finansiering av skydd, vård, forskning och övervakning av hotade arter. Resultaten av bedömningen som är genomförd av expertgrupper presenteras i 39 kapitel som omfattar olika organismgrupper.

The Red List of Finnish Species 2019

Abstract: This publication describes the fifth assessment of Finnish threatened species. The current estimated number of species in Finland is at least 48,000. Sufficient information was available for the evaluation of 22,418 species or lower taxa, which is approximately 47% of all species. In total, 2,667 species or lower taxa (11.9%) were assessed as threatened. The red-listed species, which, in addition to threatened species, include the species classified as Regionally Extinct, Near Threatened, and Data Deficient, total 6,683 (29.8%). Of the species evaluated, 489 were classified as Critically Endangered (CR), 918 as Endangered (EN), 1,260 as Vulnerable (VU), 1,912 as Near Threatened (NT), 1,792 as Data Deficient (DD), 312 as Regionally Extinct (RE), and 15,735 as of Least Concern (LC). The assessment was carried out in accordance with guidelines and criteria issued by the International Union for Conservation of Nature (IUCN) and applied in assessments at the regional level. These criteria are based on observed, estimated, inferred, or projected population size reduction; the extent of occurrence or area of occupancy; continuing decline in a small population or a very small size of population; or a quantitative analysis. In addition, species’ habitats, as well as causes of threat and threat factors, were documented in the Finnish evaluation. The majority of threatened species live in forests (31%) and in rural biotopes and other cultural habitats (24%). Of the species assessed the proportion of threatened species is highest in Alpine habitats (37.9%). The number of threatened species is highest in the hemiboreal and southern boreal zones in southern Finland. The results of the assessment are compared to the previous assessment 2010 for example with the Red List Index. In addition, proceedings of the proposals given by the steering group of the previous assessment are evaluated. The steering group of this assessment made 13 proposals for measures for the arrangement and financing of the conservation, management, research, and monitoring of threatened species. The results of the work of the expert groups responsible for the evaluation are presented in 39 organism group -specific sections.

Reference of the publication: Hyvärinen, E., Juslén, A., Kemppainen, E., Uddström, A. & Liukko, U.-M. (eds.) 2019. The 2019 Red List of Finnish Species. Ympäristöministeriö & Suomen ympäristökeskus. Helsinki. 704 p. http://hdl.handle.net/10138/299501

Publication date: 26/02/2019

10abedömning10aconservation of species10ahotade arter10aIUCN10alajiston suojelu10apunainen lista10aRed list10arödlista10askydd av arter10athreat assessment10athreatened species10auhanalaiset lajit10auhanalaisuusarviointi1 aHyvärinen, Esko1 aJuslén, Aino1 aKemppainen, Eija1 aUddström, Annika1 aLiukko, Ulla-Maija uhttp://hdl.handle.net/10138/29950100723nas a2200133 4500008004100000245004600041210004500087260001600132300000800148490000700156520033300163100001700496856007600513 2005 eng d00aMyriapods from the Eden Project, Cornwall0 aMyriapods from the Eden Project Cornwall cAutumn 2005 a1-20 v113 a

Recommended citation: Barber, T. (2005): Myriapods from the Eden Project, Cornwall. Newsletter of the British Myriapod and Isopod Group, 11: [unnumbered pages 1-2].

Bibliographic remark: Author signature is "Tony Barber". Author's name is Anthony Barber.

1 aBarber, Tony uhttps://www.bmig.org.uk/sites/www.bmig.org.uk/files/news/bmignews11.pdf00681nas a2200133 4500008004100000245008200041210006900123260001500192300000900207490000700216520022400223100001900447856008100466 2020 eng d00aPorcellionides sexfasciatus new to Britain and some rainforest goodies0 aiPorcellionides sexfasciatusi new to Britain and some rainforest c15/11/2020 a 6-70 v413 a

Recommended citation: Gregory, S. (2020): Porcellionides sexfasciatus new to Britain and some rainforest goodies. Newsletter of the British Myriapod and Isopod Group, 41: 6-7.

1 aGregory, Steve uhttps://www.bmig.org.uk/sites/www.bmig.org.uk/files/news/BMIGnews41-2020.pdf01827nas a2200241 4500008004100000022001400041245008600055210006900141260001500210300000900225490000600234520115100240653001501391653001801406653001201424653001301436653001501449653001101464100001801475700001901493700002401512856004901536 2014 eng d a1314-282800aMillipedes and centipedes in German greenhouses (Myriapoda: Diplopoda, Chilopoda)0 aMillipedes and centipedes in German greenhouses Myriapoda Diplop c11/04/2014 a1-430 v23 a

A review is given of all the literature records of millipedes and centipedes that have been found in German greenhouses together with additional records for 29 such sites. Species lists are given for 46 greenhouses investigated throughout Germany. Thirty-five diplopod and 18 chilopod species were found to occur in greenhouses, of which 15 (3 Chilopoda, 12 Diplopoda) are restricted to this type of habitat. First records for Germany include Anadenobolus monilicornis (Porat, 1876), Epinannolene cf. trinidadensis Chamberlin, 1918, Epinannolene sp., Mesoiulus gridellii Strasser, 1934, Leptogoniulus sorornus (Butler, 1876), Rhinotus purpureus (Pocock, 1894), Cryptops doriae Pocock, 1891, Lamyctes coeculus (Brölemann, 1889) and Tygarrup javanicus (Attems, 1907). The millipedes Oxidus gracilis (C. L. Koch, 1847) and Amphitomeus attemsi (Schubart, 1934) and the centipedes Lithobius forficatus (Linnaeus, 1758) and Cryptops hortensis (Donovan, 1810) are the species most frequently found in greenhouses.

10acentipedes10afirst records10aGermany10ahothouse10amillipedes10areview1 aDecker, Peter1 aReip, Hans, S.1 aVoigtländer, Karin uhttps://bdj.pensoft.net/articles.php?id=106600651nas a2200109 4500008004100000245003400041210003200075260010700107520022300214100002800437856007600465 1898 eng d00a[Letter to Ludwig Döderlein]0 aLetter to Ludwig Döderlein aStrasbourg, Grand Est, FrancebArchives de la Ville et de l’Eurométropole de Strasbourgc08/08/18983 a

Recommended citation: Verhoeff, C. W. (1898, August 8). [Letter to Ludwig Döderlein]. Archives de la Ville et de l’Eurométropole de Strasbourg, Strasbourg, Grand Est, France.

1 aVerhoeff, Carl, Wilhelm uhttps://myriatrix.myspecies.info/content/letter-ludwig-d%C3%B6derlein-100469nas a2200145 4500008004100000022001400041245005600055210005600111260001200167300000900179100002200188700003100210700002800241856005400269 1989 eng d a1021-029600aCatálogo de los artrópodos terrestres no insectos0 aCatálogo de los artrópodos terrestres no insectos c03/1989 a1-551 aMaes, Jean-Michel1 aPalacios-Vargas, José, G.1 aJiménez, María, Luisa uhttp://www.bio-nica.info/RevNicaEntomo/7-full.pdf02448nas a2200241 4500008004100000022001400041245008000055210006900135260001500204300000900219490000700228520175800235653001801993653002002011653001402031653002002045653002202065653002102087653001702108100002002125700001802145856004302163 2020 eng d a2075-445000aSoil Health and Arthropods: From Complex System to Worthwhile Investigation0 aSoil Health and Arthropods From Complex System to Worthwhile Inv c16/01/2020 a1-210 v113 a

The dramatic increase in soil degradation in the last few decades has led to the need to identify methods to define not only soil quality but also, in a holistic approach, soil health. In the past twenty years, indices based on living communities have been proposed alongside the already proven physical-chemical methods. Among them, some soil invertebrates have been included in monitoring programs as bioindicators of soil quality. Being an important portion of soil fauna, soil arthropods are involved in many soil processes such as organic matter decomposition and translocation, nutrient cycling, microflora activity regulation and bioturbation. Many studies have reported the use of soil arthropods to define soil quality; among taxa, some have been explored more in depth, typically Acari and Collembola, while generally less abundant groups, such as Palpigradi or Embioptera, have not been investigated much. This paper aims to evaluate and compare the use of different soil microarthropod taxa in soil degradation/quality studies to highlight which groups are the most reported for soil monitoring and which are the most sensitive to soil degradation. We have decided not to include the two most present and abundant taxa, Acari and Collembola, in this paper in consideration of the vast amount of existing literature and focus the discussion on the other microarthropod groups. We reported some studies for each taxon highlighting the use of the group as soil quality indicator. A brief section reporting some indices based on soil microarthropods is proposed at the end of this specific discussion. This paper can be considered as a reference point in the use of soil arthropods to estimate soil quality and health.

10abioindicators10amacroarthropods10amesofauna10amicroarthropods10asoil biodiversity10asoil degradation10asoil quality1 aMenta, Cristina1 aRemelli, Sara uhttps://www.mdpi.com/2075-4450/11/1/5401282nas a2200145 4500008004100000022001400041245009000055210006900145260001200214300001000226520079400236100002701030700002001057856005901077 2020 eng d a0242-177100aMyriapodes chilopodes et diplopodes des souterrains de Paris et de sa proche banlieue0 aMyriapodes chilopodes et diplopodes des souterrains de Paris et  c03/2020 a39-473 a

Une campagne d’échantillonnage des invertébrés a été menée dans les réseaux souterrains de la ville de Paris et de la banlieue proche. Le présent travail présente les résultats obtenus concernant les myriapodes (chilopodes et diplopodes) issus de ces récoltes. Six espèces de chilopodes (Classe Chilopoda) et neuf espèces de diplopodes (Classe Diplopoda) ont pu être identifiées. Parmi ces dernières, certaines espèces sont nouvelles pour les départements des Yvelines, des Hauts-de-Seine et de la Seine-Saint-Denis. Ces résultats contribuent à l'enrichissement des inventaires et atlas des myriapodes de France (project INAMYFRA).

1 aGeoffroy, Jean-Jacques1 aFerrand, Marina uhttps://publications.ffspeleo.fr/bulletin.php?id=2599900710nas a2200097 4500008004100000245007300041210006900114520032800183100002000511856008100531 2020 eng d00aNewsletter of the British Myriapod and Isopod Group, 41, Autumn 20200 aNewsletter of the British Myriapod and Isopod Group 41 Autumn 203 a

The newsletter includes items on: Centipede atlas update. The Darwin Tree of Life project. Luminous centipedes. Some rainforest goodies. Paul Richards giving an introduction to millipedes. Hebridean fossil millipedes in the Times.

Published: 15.11.2020 (website upload date, communicated by Steve Gregory).

1 aRead, Helen, J. uhttps://www.bmig.org.uk/sites/www.bmig.org.uk/files/news/BMIGnews41-2020.pdf03159nas a2200313 4500008004100000022001400041245007400055210006900129260001500198300000900213490000700222520226400229100001202493700001702505700001502522700002102537700001502558700002502573700001402598700001802612700002502630700001802655700002002673700002302693700001902716700002302735700003402758856005302792 2020 eng d a1544-917300aMillipede genomes reveal unique adaptations during myriapod evolution0 aMillipede genomes reveal unique adaptations during myriapod evol c29/09/2020 a1-240 v183 a

The Myriapoda, composed of millipedes and centipedes, is a fascinating but poorly understood branch of life, including species with a highly unusual body plan and a range of unique adaptations to their environment. Here, we sequenced and assembled 2 chromosomal-level genomes of the millipedes Helicorthomorpha holstii (assembly size = 182 Mb; shortest scaffold/contig length needed to cover 50% of the genome [N50] = 18.11 Mb mainly on 8 pseudomolecules) and Trigoniulus corallinus (assembly size = 449 Mb, N50 = 26.78 Mb mainly on 17 pseudomolecules). Unique genomic features, patterns of gene regulation, and defence systems in millipedes, not observed in other arthropods, are revealed. Both repeat content and intron size are major contributors to the observed differences in millipede genome size. Tight Hox and the first loose ecdysozoan ParaHox homeobox clusters are identified, and a myriapod-specific genomic rearrangement including Hox3 is also observed. The Argonaute (AGO) proteins for loading small RNAs are duplicated in both millipedes, but unlike in insects, an AGO duplicate has become a pseudogene. Evidence of post-transcriptional modification in small RNAs—including species-specific microRNA arm switching—providing differential gene regulation is also obtained. Millipedes possesses a unique ozadene defensive gland unlike the venomous forcipules found in centipedes. We identify sets of genes associated with the ozadene that play roles in chemical defence as well as antimicrobial activity. Macro-synteny analyses revealed highly conserved genomic blocks between the 2 millipedes and deuterostomes. Collectively, our analyses of millipede genomes reveal that a series of unique adaptations have occurred in this major lineage of arthropod diversity. The 2 high-quality millipede genomes provided here shed new light on the conserved and lineage-specific features of millipedes and centipedes. These findings demonstrate the importance of the consideration of both centipede and millipede genomes—and in particular the reconstruction of the myriapod ancestral situation—for future research to improve understanding of arthropod evolution, and animal evolutionary genomics more widely.

1 aQu, Zhe1 aNong, Wenyan1 aSo, Wai, L1 aBarton-Owen, Tom1 aLi, Yiqian1 aLeung, Thomas, C. N.1 aLi, Chade1 aBaril, Tobias1 aWong, Annette, Y. P.1 aSwale, Thomas1 aChan, Ting-Fung1 aHayward, Alexander1 aNgai, Sai-Ming1 aHui, Jerome, H. L.1 aFernandez-Valverde, Selene, L uhttps://dx.plos.org/10.1371/journal.pbio.300063601302nas a2200229 4500008004100000022001400041245009000055210006900145260001500214300001000229490000700239520063800246653001000884653001400894653001700908653001100925653002300936653001600959100002700975700002201002856004801024 2020 eng d a1768-144800aTerrestrial invertebrate fauna of Polish caves – a summary of 100 years of research0 aTerrestrial invertebrate fauna of Polish caves a summary of 100  c13/02/2020 a45-690 v333 a

The year 2018 is particularly important in the history of zoological research in Poland. A hundred years ago, Kazimierz Demel published the first work concerning the terrestrial cave fauna of caves in the Ojców area. In this paper we present the extent of research on the terrestrial invertebrate fauna of Polish caves in the last 100 years. All accessible research papers that have been published during this period were analysed. Based on published literature, 593 species of terrestrial invertebrate were recorded in Polish caves. Additionally, detailed list of species of individual taxonomic groups was provided.

10acaves10achecklist10ainvertebrate10aPoland10asubterranean fauna10aterrestrial1 aKocot-Zalewska, Joanna1 aDomagała, Paweł uhttps://subtbiol.pensoft.net/article/48805/01305nam a2200145 4500008004100000050001400041245012200055210006900177260004300246300001400289490000600303520076800309100002301077856005901100 1847 eng d aQL449.K6500aSystem der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden0 aSystem der Myriapoden mit den Verzeichnissen und Berichtigungen  aRegensburg, Germanyb Friedrich Pustet aiii + 2720 v33 a

Recommended citation: Koch, C. L. (1847): System der Myriapoden mit den Verzeichnissen und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden. Hefte 1-40. Friedrich Pustet, Regensburg, iii + 272 pp. + 10 pls. https://doi.org/10.5962/bhl.title.49866. In: Panzer, G. W. F. and G. A. W. Herrich‐Schäffer (Eds.). Kritische Revision der Insectenfaune Deutschlands. Hefte 1-190. III. Bändchen, enthaltend die Verzeichnisse und Berichtigungen zu Deutschlands Crustaceen, Myriapoden und Arachniden und ein System der Myriapoden von C. L. Koch.

Bibliographic remark: Herrich‐Schäffer is the sole editor of the Koch’s series. See Nagel & Grieder (2019: 36).

1 aKoch, Carl, Ludwig uhttps://www.biodiversitylibrary.org/bibliography/4986602624nas a2200229 4500008004100000022001400041245006100055210006100116260001500177300000900192490000800201520194600209653001902155653002702174653001802201653002302219653001902242100002002261700002402281700002202305856006702327 2020 eng d a0965-174800aDiversity and evolution of the P450 family in arthropods0 aDiversity and evolution of the P450 family in arthropods c22/10/2020 a1-220 v1273 a

The P450 family (CYP genes) of arthropods encodes diverse enzymes involved in the metabolism of foreign compounds and in essential endocrine or ecophysiological functions. The P450 sequences (CYPome) from 40 arthropod species were manually curated, including 31 complete CYPomes, and a maximum likelihood phylogeny of nearly 3000 sequences is presented. Arthropod CYPomes are assembled from members of six CYP clans of variable size, the CYP2, CYP3, CYP4 and mitochondrial clans, as well as the CYP20 and CYP16 clans that are not found in Neoptera. CYPome sizes vary from two dozen genes in some parasitic species to over 200 in species as diverse as collembolans or ticks. CYPomes are comprised of few CYP families with many genes and many CYP families with few genes, and this distribution is the result of dynamic birth and death processes. Lineage-specific expansions or blooms are found throughout the phylogeny and often result in genomic clusters that appear to form a reservoir of catalytic diversity maintained as heritable units. Among the many P450s with physiological functions, six CYP families are involved in ecdysteroid metabolism. However, five so-called Halloween genes are not universally represented and do not constitute the unique pathway of ecdysteroid biosynthesis. The diversity of arthropod CYPomes has only partially been uncovered to date and many P450s with physiological functions regulating the synthesis and degradation of endogenous signal molecules (including ecdysteroids) and semiochemicals (including pheromones and defense chemicals) remain to be discovered. Sequence diversity of arthropod P450s is extreme, and P450 sequences lacking the universally conserved Cys ligand to the heme have evolved several times. A better understanding of P450 evolution is needed to discern the relative contributions of stochastic processes and adaptive processes in shaping the size and diversity of CYPomes.

10aDetoxification10aEcdysteroid metabolism10aGene clusters10aOrthologous groups10aP450 Evolution1 aDermauw, Wannes1 aVan Leeuwen, Thomas1 aFeyereisen, René uhttps://linkinghub.elsevier.com/retrieve/pii/S096517482030179X00403nas a2200121 4500008004100000245003300041210003300074260007300107300001200180490000600192100002900198856005400227 1887 eng d00aNew North American myriapods0 aNew North American myriapods b[Lawrence, Kan., etc.]New York Entomological Society [etc.]c08/1887 a81–830 v31 aBollman, Charles, Harvey uhttps://www.biodiversitylibrary.org/page/1140597100536nas a2200133 4500008004100000245006900041210006900110260006600179300001400245490000700259520005600266100002900322856005100351 1888 eng d00aDescriptions of fourteen new species of North American myriapods0 aDescriptions of fourteen new species of North American myriapods bWashington :Smithsonian Institution Press, [etc.]c19/09/1888 a617–6270 v103 a

Remark: Published in signatures 39 and 40.

1 aBollman, Charles, Harvey uhttps://www.biodiversitylibrary.org/part/23758