02194nas a2200217 4500008004100000022001400041245016500055210006900220260001200289300001400301490000700315520143900322653002601761653001401787653001301801653001701814653002601831100002301857700002001880856007601900 2018 eng d a1467-803900aA first phylogenetic analysis of the pill millipedes of the order Glomerida, with a special assessment of mandible characters (Myriapoda, Diplopoda, Pentazonia)0 afirst phylogenetic analysis of the pill millipedes of the order  c03/2018 a214 - 2280 v473 a

The pill millipedes of the order Glomerida are a moderately diverse group with a classical Holarctic distribution pattern. Their classification is based on a typological system utilizing mainly a single character complex, the male telopods. In order to infer the apomorphies of the Glomerida, to elucidate its position in the Pentazonia, and to test the monophyly of its families and subfamilies, we conduct the first phylogenetic analysis of the order. To provide additional characters, we comparatively analyze the mandible using scanning electron microscopy. The final character matrix consists of 69 characters (11 mandible characters) and incorporates 22 species from 20 of the 34 pill millipede genera, representing all families and subfamilies, except the monotypic Mauriesiinae. Two species from each of the two other Pentazonian orders Sphaerotheriida and Glomeridesmida, as well as two Spirobolida, are included as outgroup taxa. The Glomerida are recovered as monophyletic and are supported by five apomorphies. Within the Pentazonia, the Glomeridesmida are recovered as the sister group to the classical Oniscomorpha (Sphaerotheriida + Glomerida) with weak support. The analysis provides little resolution within the Glomerida, resulting in numerous polytomies. Further morphological characters and/or the addition of molecular analyses are needed to produce a robust phylogenetic classification of the Glomerida.

10aapomorphic characters10aGlomerida10amandible10aOniscomorpha10aphylogenetic analysis1 aOeyen, Jan, Philip1 aWesener, Thomas uhttps://www.sciencedirect.com/science/article/abs/pii/S146780391830020302208nas a2200229 4500008004100000022001400041245019300055210006900248260001400317300001200331490000800343520145200351653000801803653001401811653001601825653001801841653001101859653001301870100002301883700002001906856005201926 2015 eng d a1313-298900aSteps towards a phylogeny of the pill millipedes: non-monophyly of the family Protoglomeridae, with an integrative redescription of Eupeyerimhoffia archimedis (Diplopoda, Glomerida)0 aSteps towards a phylogeny of the pill millipedes nonmonophyly of c3006/2015 a49 - 640 v5103 a

Eupeyerimhoffia archimedis (Strasser, 1965) is redescribed based on several specimens collected at a number of sites close to the type locality on Sicily, Italy. Scanning electron microscopy is used to illustrate several unusual morphological characters for a member of the Glomerida for the first time. A fragment of the mitochondrial COI gene (668bp) is sequenced for the first time in Eupeyerimhoffia to provide a species-specific barcode and to gain first insights into the genetic distances between the genera in the widespread family Protoglomeridae. The novel sequences are compared to representatives of all other genera of the family: Protoglomeris vasconica (Brölemann, 1897) from northern Spain, the dwarfed Glomerellina laurae Silvestri, 1908 from Italy and Glomeroides primus (Silvestri, 1929) from western North America. The addition of COI sequences from the two other families of the Glomerida renders the family Protoglomeridae paraphyletic with Glomeroides primus being more closely related to Glomeridella minima (Latzel, 1884) than to the other genera in the family. The large genetic distances (13.2–16.8%) between Eupeyerimhoffia and the other genera in the order, as well as its unusual morphological characters, including unique morphological adaptations to roll into a ball, are probably an indication of the old age of the group.

10aCOI10aGlomerida10aintegrative10aredescription10aSicily10aTaxonomy1 aOeyen, Jan, Philip1 aWesener, Thomas uhttp://zookeys.pensoft.net/articles.php?id=5524