02982nas a2200265 4500008004100000022002500041245018800066210006900254260001500323300001000338490000800348520201400356653001402370653001702384653001402401653001602415653002202431653001602453100001902469700002702488700002402515700002202539700001802561856013702579 2016 eng d a1313-2989, 1313-297000aA taxonomic review of the centipede genus Scolopendra Linnaeus, 1758 (Scolopendromorpha, Scolopendridae) in mainland Southeast Asia, with description of a new species from Laos0 ataxonomic review of the centipede genus iScolopendrai Linnaeus 1 c17/05/2016 a1-1240 v5903 a

The centipede genus Scolopendra in mainland Southeast Asia is reviewed taxonomically based on morphological characters, informed by a molecular phylogenetic analysis using sequences from three mitochondrial and nuclear genes (COI, 16S rRNA and 28S rRNA). Eight nominal species of Scolopendra, namely S. morsitans Linnaeus, 1758, S. subspinipes Leach, 1816, S. dehaani Brandt, 1840, S. multidens Newport, 1844, S. calcarata Porat, 1876, S. japonica Koch, 1878, S. pinguis Pocock, 1891, and S. dawydoffi Kronmüller, 2012, are redescribed together with some revision of type materials. Geographical variation in each species has been compiled with reference to samples that span their distribution ranges in Southeast Asia and some parts of neighbouring areas such as East Asia, the Indian Ocean, and Africa. Comparative study of traditional taxonomic characters from external morphology provides further information to distinguish some closely related species. Scolopendra cataracta Siriwut, Edgecombe & Panha, sp. n., is described from the southern part of Laos, with additional records in Thailand and Vietnam. The phylogenetic framework for Southeast Asian Scolopendra recognizes S. calcarata + S. pinguis, S. morsitans, and a S. subspinipes group that unites the other six species as the main clades. Within the S. subspinipes group, two monophyletic groups can be distinguished by having either slender or short, thick ultimate leg prefemora and different numbers of apical spines on the coxopleuron. Scolopendra arborea Lewis, 1982, is placed in subjective synonymy with S. dehaani. A survey of external morphology of the genital segments confirms its potential for improving species identification in Scolopendra. Some observations on biology and behaviour are recorded based on field surveys in this area.

10aChilopoda10adistribution10aphylogeny10aScolopendra10aspecies diversity10asystematics1 aSiriwut, Warut1 aEdgecombe, Gregory, D.1 aSutcharit, Chirasak1 aTongkerd, Piyoros1 aPanha, Somsak uhttps://myriatrix.myspecies.info/content/taxonomic-review-centipede-genus-scolopendra-linnaeus-1758-scolopendromorpha-scolopendridae02692nas a2200229 4500008004100000022002500041245014800066210006900214260001500283300001200298490000700310520196600317653000802283653001202291653001402303653001702317100002402334700002002358700001802378700002302396856004302419 2022 eng d a1860-0743, 1435-193500aA new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species0 anew genus of Pseudospirobolellidae Diplopoda Spirobolida from li c19/08/2022 a313-3260 v983 a

A new genus of the millipede family Pseudospirobolellidae, Siliquobolellus gen. nov., is described from limestone mountains in Thailand, based on three new species, viz. Siliquobolellus amicusdraconis gen. et sp. nov. from Uthaithani Province, Siliquobolellus constrictus gen. et sp. nov. from Prachuap Khiri Khan Province and Siliquobolellus prasankokae gen. et sp. nov. (type species) from Lampang Province. The descriptions are based on gonopod morphology and mitochondrial DNA data (COI barcodes). The COI barcodes grouped the three new species in a well-supported Siliquobolellus gen. nov. clade. The mean interspecific COI sequence divergence among the three new species was 12% (range: 8–15%). The mean intergeneric COI sequence divergence between Siliquobolellus gen. nov., Coxobolellus Pimvichai, Enghoff, Panha & Backeljau, 2020, and Pseudospirobolellus Carl, 1912 was 19% (range: 14–23%). Three conspicuous gonopodal synapomorphies differentiate Siliquobolellus gen. nov. from other pseudospirobollellid genera: (1) the telopodital part of the posterior gonopod forms a deep concavity, (2) the telopodite of the anterior gonopod is directed distad and does not reach the tip of the coxal part of the anterior gonopod, and (3) the tip of the anterior gonopod coxa is narrowed, curving mesad. As such, the monophyly of the new genus is well supported by both morphological and mitochondrial DNA data. A distribution map and an identification key to the species are provided.

10aCOI10aGonopod10amonophyly10asynapomorphy1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://zse.pensoft.net/article/90032/02428nas a2200277 4500008004100000022002500041245023700066210006900303260001500372300001100387490000700398520148100405653001801886653001301904653002201917653002101939653001401960653002401974653001301998653001802011653001202029100002402041700001802065700002302083856004402106 2022 eng d a1447-2600, 1445-522600aCombining mitochondrial DNA and morphological data to delineate four new millipede species and provisional assignment to the genus Apeuthes Hoffman & Keeton (Diplopoda : Spirobolida : Pachybolidae : Trigoniulinae)0 aCombining mitochondrial DNA and morphological data to delineate  c16/02/2022 a91-1120 v363 a

Hitherto, the millipede genus Apeuthes (family Pachybolidae, subfamily Trigoniulinae) was only known from three species described in Vietnam based on morphological characters. The present study uses two partial mitochondrial gene fragments (cytochrome c oxidase I (COI) and 16S ribosomal RNA) and morphology to define four new Apeuthes species from Malaysia, Thailand and Vietnam: A. fimbriatus, sp. nov., A. longeligulatus, sp. nov., A. pollex, sp. nov. and ?A. spininavis, sp. nov. The intraspecific COI sequence divergence of two Apeuthes species is 3–7% (mean: 5%) and the interspecific divergence of five species is 11–16% (mean: 13.7%). All members of the genus share unique male characters, viz the posterior gonopod telopodite with several dentate, serrate or tuberculate lamellae in a boat-like cavity or a boat-like cavity covered with spines. The delimitation of the four new species is supported by the congruence between mitochondrial DNA and morphological data. However, while the monophyly of Trigoniulinae is well supported, the relationships within this subfamily, and particularly among Apeuthes species, including the monophyly of Apeuthes, lack strong support. Therefore assignment of the four new species, and particularly of ?A. spininavis sp. nov., to the genus Apeuthes is tentative and awaits a comprehensive revision of the group.

10aDNA barcoding10aMalaysia10amitochondrial DNA10amonotypic genera10aphylogeny10aspecies delineation10aThailand10aTrigoniulinae10aVietnam1 aPimvichai, Piyatida1 aPanha, Somsak1 aBackeljau, Thierry uhttps://www.publish.csiro.au/IS/IS2103803135nas a2200241 4500008004100000022001400041245025000055210006900305260001500374300001200389490000700401520225900408653002202667653001602689653001402705653002002719653002502739100002402764700002002788700001802808700002302826856004402849 2018 eng d a1445-522600aMorphological and mitochondrial DNA data reshuffle the taxonomy of the genera Atopochetus Attems, Litostrophus Chamberlin and Tonkinbolus Verhoeff (Diplopoda: Spirobolida: Pachybolidae), with descriptions of nine new species0 aMorphological and mitochondrial DNA data reshuffle the taxonomy  c01/02/2018 a159-1950 v323 a

Species-level taxonomy and phylogeny of two genera of South-East (SE) Asian pachybolid millipedes are analysed with a combination of morphological characters and DNA sequences (two mitochondrial gene fragments: COI and 16S rRNA). Strong support is found for the genera Litostrophus Chamberlin, 1921 and Atopochetus Attems, 1953 and for a clade consisting of Litostrophus + Atopochetus. Four species of Litostrophus are recognised and (re)described: L. segregatus Chamberlin, 1921 (type species of the genus), L. scaber (Verhoeff, 1938), comb. nov., L. chamaeleon, sp. nov. and L. saraburensis, sp. nov. The genus Tonkinbolus Verhoeff, 1938 (type species T. scaber Verhoeff, 1938) is synonymised under Litostrophus. Atopochetus (type species A. rubropunctatus Attems, 1953), hitherto considered a dubious synonym of Aulacobolus Pocock, 1903, is re-instated for several species until now placed in Tonkinbolus and seven new species. All in all, 10 species of Atopochetus are (re)described: A. dollfusii (Pocock, 1893) (= Aulacobolus rubropunctatus Attems, 1938, syn. nov.), A. moulmeinensis (Pocock, 1893), A. sumatranus (Carl, 1906) (= Trachelomegalus laciniatus Attems, 1937, syn. nov.), A. anaticeps, sp. nov., A. helix, sp. nov., A. setiferus, sp. nov., A. spinimargo, sp. nov., A. truncatus, sp. nov., A. uncinatus, sp. nov. and A. weseneri, sp. nov. Three species recently included in Tonkinbolus are left incertae sedis because they are based exclusively on female type specimens: Spirobolus capucinus Porat, 1896, S. caudulanus Karsch, 1881 and S. macrurus Pocock, 1893. The results of the DNA analysis (COI and 16S rRNA) are congruent with morphological (gonopodal) characters in terms of delimitation of species of Litostrophus and Atopochetus. This is the first DNA study of SE Asian Pachybolidae and as such it provides a basis for further evolutionary and biogeographic studies of SE Asian millipedes.

10amitochondrial DNA10anew species10aphylogeny10aSouth-East Asia10aspecies delimitation1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://www.publish.csiro.au/is/IS1705201621nas a2200241 4500008004100000022001400041245021400055210006900269260001500338300000900353490000800362520077600370653001801146653001501164653001601179653001401195653001301209100002401222700002001246700001801266700002301284856007201307 2016 eng d a2118-977300aA revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part V Nine new spe c24/05/2016 a1-370 v1993 a

The Thyropygus opinatus subgroup (Diplopoda: Harpagophoridae) of the T. allevatus group in Thailand is revised. Based on a phylogenetic analysis of mtDNA sequence data, it is merged with the T. bifurcus subgroup to form an extended T. opinatus subgroup. Nine new species are described: Thyropygus cimi sp. nov. and T. forceps sp. nov. from Nakhonsrithammarat Province, T. culter sp. nov., T. planispina sp. nov., T. undulatus sp. nov. and T. ursus sp. nov. from Krabi Province, T. mesocristatus sp. nov. from Songkhla Province, T. navychula sp. nov. from Phang-Nga Province and T. sutchariti sp. nov. from Phetchaburi Province.

10aDNA barcoding10amillipedes10anew species10aphylogeny10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/32202099nas a2200241 4500008004100000022001400041245011800055210006900173260001500242300001200257490000700269520133500276653001301611653002701624653002201651653001401673653001901687653001301706100002401719700002001743700001801763856007601781 2013 eng d a1095-951300aMolecular phylogeny of the Thyropygus allevatus group of giant millipedes and some closely related groups.0 aMolecular phylogeny of the iThyropygus allevatusi group of giant c20/11/2013 a170-1830 v713 a

Giant cylindrical millipedes of the family Harpagophoridae, especially species of the genus Thyropygus, are broadly distributed in Thailand and nearby countries. They show a great deal of variation in body size, color patterns and gonopodal characters. Phylogenetic analyses of 26 nominate species from six genera in the subfamilies Harpagophorinae and Rhynchoproctinae, as well as nine new morphotypes (regarded as new species), were performed with the DNA sequences from two mitochondrial gene fragments (16S rRNA and COI). The genus Thyropygus (Harpagophorinae) was recovered as monophyletic under all analyses, whilst the representatives of Rhynchoproctinae also formed a monophyletic group. However, the analyses suggested that the tribe Gonoplectini should be elevated to a separate subfamily, Gonoplectinae. The molecular analyses were largely (but not totally) congruent with, and so supported the usefulness of, gonopodal characters for the classification and identification of harpagophorid millipedes, and additionally supported previous studies on the delimitation of species and subgroups. This is the first molecular study inside the family Harpagophoridae and provides the basis for further studies of the evolutionary processes and biogeographic patterns of millipedes in Southeast Asia.

10aendemism10aGeographical isolation10amitochondrial DNA10aphylogeny10aSoutheast Asia10asympatry1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.sciencedirect.com/science/article/abs/pii/S105579031300418101673nas a2200265 4500008004100000022002500041245017100066210006900237260001500306300001000321490000900331520080000340653001301140653001501153653001401168653001601182653001301198100002201211700002401233700002401257700002001281700002701301700001801328856006101346 2012 eng d a1175-5334, 1175-532600aRevision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae)0 aRevision of the rare centipede genus iSterropristesi Attems 1934 c14/09/2012 a35-520 v34843 a

The genus Sterropristes Attems, 1934, currently classified in the scolopendrid subfamily Otostigminae and tribe Sterropristini, is revised. The monotypic genus Malaccolabis Verhoeff, 1937, is synonymized with Sterropristes. Sterropristes sarasinorum Attems, 1934 from Tomohon, Celebes, Indonesia (type species of Sterropristes) and S. metallicus (Verhoeff, 1937) from Penang Hill, Penang, Malaysia (type species of Malaccolabis) are redescribed based on their type material, together with newly collected topotypes of S. metallicus. Sterropristes violaceus Muadsub and Panha n. sp., found under leaves on islands in the Andaman Sea, south-western Thailand, is described.

10aMalaysia10amorphology10aMyriapoda10asystematics10aTaxonomy1 aMuadsub, Sawitree1 aSutcharit, Chirasak1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aEdgecombe, Gregory, D.1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.3484.1.201130nas a2200217 4500008004100000022002500041245014400066210006900210260001500279300001000294490000900304520042000313653001400733653001600747653001300763653001300776100002400789700002000813700001800833856006100851 2011 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 4 the iT cuisi c29/07/2011 a37-480 v29803 a

The Thyropygus cuisinieri subgroup of the T. allevatus group is revised. A new species is described from Thailand: T. jarukchusri n. sp. The following species are re-described: T. cuisinieri Carl, 1917, T. carli Attems, 1938, and T. foliaceus (Demange, 1961), new status (elevated from subspecies status under T. cuisinieri).

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2980.1.301618nas a2200229 4500008004100000022002500041245014300066210006900209260001500278300001000293490000900303520087700312653001401189653001601203653002001219653001301239653001301252100002401265700002001289700001801309856006101327 2011 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 3 the iT indur c06/07/2011 a47-680 v29413 a

The Thyropygus induratus subgroup of the T. allevatus group is revised. Three new species are described from Thailand: T. dormiens n. sp., T. laterolobatus n. sp. and T. macrosiamensis n. sp. The following species are redescribed and re-elevated from subspecies status under T. allevatus (Karsch, 1881): Thyropygus induratus Attems, 1936 (= T. punctatus Attems, 1938, new synonym), Thyropygus quietus Attems, 1938, Thyropygus resimus Attems, 1938, and Thyropygus uncinatus (Demange, 1961). Thyropygus siamensis Verhoeff, 1938 (= Thyropisthus ligulus Demange, 1961, new synonym), is re-described. A case of species with identical gonopods but very different body size (T. induratus and T. quietus) is discussed.

10amillipede10anew species10asize difference10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2941.1.301583nas a2200169 4500008004100000022001400041245023000055210006900285260001500354300001000369490000700379520092100386100002401307700002001331700001801351856004401369 2010 eng d a1445-522600aThe Rhynchoproctinae, a south-east Asiatic subfamily of giant millipedes: cladistic analysis, classification, four new genera and a deviating new species from north-west Thailand (Diplopoda : Spirostreptida : Harpagophoridae)0 aRhynchoproctinae a southeast Asiatic subfamily of giant milliped c17/05/2010 a51-800 v243 a

The subfamily Rhynchoproctinae is revised at the generic level. Four new genera and a remarkable new species are described: Alienostreptus, gen. nov. from Vietnam, Armatostreptus, gen. nov., Heptischius, gen. nov., and Heptischius lactuca, sp. nov. from Thailand, and Prominulostreptus, gen. nov. from China. Agaricogonopus Zhang & Zhang, 1997, is removed from synonymy with Junceustreptus Demange, 1961. All 14 genera of the subfamily share three diagnostic gonopodal characters: the posterior surface of the anterior coxal fold forms a deep concavity, there is a mesal flap on the gonopod coxa, and the posterior coxal fold is very low. A cladistic analysis based on morphological data supports monophyly of Rhynchoproctinae and the distincness of the newly described genera. A dichotomous key to the genera is included.

1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.publish.csiro.au/is/is0905201206nas a2200217 4500008004100000022002500041245014200066210006900208260001500277300000900292490000900301520049900310653001400809653001600823653001300839653001300852100002400865700002000889700001800909856006100927 2009 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 2: the T. bifurcus subgroup (Diplopoda, Spirostreptida, Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 2 the iT bifur c22/07/2009 a1-150 v21653 a

The Thyropygus bifurcus subgroup of the T. allevatus group is revised. Four new species are described from Thailand: T. demangei n. sp., T. quadricuspis n. sp., T. richardhoffmani n. sp., from Trang province and T. casjeekeli n. sp., from Krabi province. The other species of the T. bifurcus subgroup, viz., T. bifurcus (Demange, 1986), and T. enghoffi (Demange, 1989), are redescribed.

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2165.1.101898nas a2200217 4500008004100000022002500041245014200066210006900208260001500277300001000292490000900302520119000311653001401501653001601515653001301531653001301544100002401557700002001581700001801601856006101619 2009 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 1 the iT opina c20/02/2009 a17-500 v20163 a

The Thyropygus opinatus subgroup of the T. allevatus group is revised. The T. opinatus subgroup corresponds to the genus Cornugonus Demange, 1961, which is formally synonymized under Thyropygus. Eight new species are described from Thailand: T. bearti n. sp., T. brachyacanthus n. sp., T. loxia n. sp., from Suratthani province, T. bispinus n. sp., from Uthaithani and Phrae provinces, T. bispinispatula n. sp., from Chumphon province, T. chelatus n. sp., from Nakhonsrithammarat province, T. cristagalli n. sp., from Phang Nga province and T. erectus n. sp., from Satun province. The other species of the T. opinatus subgroup, viz., T. opinatus (Karsch, 1881), n. comb., T. floweri (Demange, 1961), n. comb., T. implicatus (Demange, 1961), n. comb., and T. inflexus (Demange, 1989), n. comb., are redescribed. New records are given for T. opinatus, T. implicatus and T. floweri. The type locality of T. floweri is shown to be in Thailand rather than Malaysia.

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2016.1.201551nas a2200229 4500008004100000022002500041245012300066210006900189260001500258300000900273490000900282520082100291653001001112653001001122653001401132653001301146100002701159700002501186700002601211700001801237856006601255 2017 eng d a1175-5334, 1175-532600aA checklist of the millipedes (Diplopoda) of Myanmar, with an updated list of Leonardo Fea’s collecting localities 0 achecklist of the millipedes Diplopoda of Myanmar with an updated c16/11/2017 a1-460 v43503 a

At present, the millipede fauna of Myanmar comprises 92 species from 34 genera, 13 families and 8 orders, mostly described in 1889–1896. All literature records are cited with updates on species identities, as well as numerous taxonomic problems which make the number of species and even genera still imprecise. The Myanmar millipede fauna contains 70 endemic and five widespread synanthropic species. One species is found to have erroneously been recorded from Myanmar, and is ejected from the list of Myanmar millipedes also because of its uncertain taxonomic status. A complete bibliography on the millipedes of Myanmar, an updated list of the collecting localities and a map of the journeys of Leonardo Fea, the principal collector of Diplopoda in Burma, are also presented. 

10aBurma10afauna10amillipede10aTaxonomy1 aLikhitrakarn, Natdanai1 aJirapatrasilp, Parin1 aGolovatch, Sergei, I.1 aPanha, Somsak uhttps://www.biotaxa.org/Zootaxa/article/view/zootaxa.4350.1.100785nas a2200169 4500008004100000245018800041210006900229260001200298100002900310700001800339700003300357700003000390700002000420700002000440700002500460856013000485 2012 eng d00aDaily Activities of the Giant Pill-Millipede Zephronia cf. viridescens Attems, 1936 (Diplopoda: Sphaerotheriida: Zephroniidae) in a Deciduous Forest in Northern Thailand0 aDaily Activities of the Giant PillMillipede iZephroniai cf iviri c01/20121 aWongthamwanich, Nattarin1 aPanha, Somsak1 aSitthicharoenchai, Duangkhae1 aPradatsundarasar, Art-Ong1 aSeelanan, Tosak1 aEnghoff, Henrik1 aThirakhupt, Kumthorn uhttps://myriatrix.myspecies.info/content/daily-activities-giant-pill-millipede-zephronia-cf-viridescens-attems-1936-diplopoda01236nas a2200229 4500008004100000022001300041245014900054210006900203260001500272300001100287490000700298520047600305653001700781653001400798653001400812653001200826653001600838100002500854700001400879700001800893856009500911 2020 eng d a021-244500aA revision of dragon millipedes. V. The new genus Burmaxytes for two new species from Myanmar (Diplopoda: Polydesmida: Paradoxosomatidae)0 arevision of dragon millipedes V The new genus iBurmaxytesi for t c31/03/2020 a88-1000 v353 a

A new genus of dragon millipedes, Burmaxytes, is described from Myanmar, based on two new species: B. mombergi (type species) from Kayin State, and B. whitteni from Mon State. The new genus, constituting a sixth group of dragon millipedes, is discussed, and a distribution map is provided. The two new species were exclusively found living on humid limestone rock walls, and are endemic and restricted to limestone habitats in Myanmar.

10abiodiversity10alimestone10amillipede10aMyanmar10anew species1 aSrisonchai, Ruttapon1 aLin, Aung1 aPanha, Somsak uhttps://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/app/uploads/2020/03/s35rbz88-100.pdf02849nas a2200229 4500008004100000022001400041245017100055210006900226260001500295300001200310490000700322520208700329653002202416653001402438653001402452653002402466100002402490700002002514700001802534700002302552856004402575 2020 eng d a1445-522600aIntegrative taxonomy of the new millipede genus Coxobolellus, gen. nov. (Diplopoda : Spirobolida : Pseudospirobolellidae), with descriptions of ten new species0 aIntegrative taxonomy of the new millipede genus iCoxobolellusi g c14/08/2020 a591-6170 v343 a

Pseudospirobolellidae is a poorly known family of spirobolidan millipedes with only two genera and five described species. Yet, the descriptive taxonomy and molecular systematics of this group have been largely neglected. Therefore, the present work presents an integrative taxonomic study of new pseudospirobolellid taxa in Thailand. To this end, two mitochondrial gene fragments (COI and 16S rRNA) combined with morphological characters were used to define the genus Coxobolellus, gen. nov. with 10 new species, viz. C. albiceps, sp. nov., C. compactogonus, sp. nov., C. fuscus, sp. nov., C. nodosus, sp. nov., C. serratus, sp. nov., C. simplex, sp. nov., C. tenebris, sp. nov., C. tigris, sp. nov., C. transversalis, sp. nov. and C. valvatus, sp. nov. The interspecific COI sequence divergences among the new species ranged from 6 to 15%. The intergeneric COI sequence divergence between species of Coxobolellus, gen. nov., Benoitolus birgitae and Pseudospirobolellus sp. ranged from 20 to 23%. Three major morphological differences separate Coxobolellus, gen. nov. from Benoitolus and Pseudospirobolellus, namely (1) the protruding process on the 3rd (and 4th) coxae on male legs, (2) the posterior gonopod telopodite divided into two parts, and (3) a conspicuous opening pore at the mesal margin at the end of the coxal part of the posterior gonopod. Thus, the new genus is well supported by both mtDNA and morphological evidence, while the delimitation of the 10 new species is supported by the congruence between mtDNA and morphological data. Yet, with respect to the relationships of Benoitolus birgitae, morphological data suggest a similarity with Coxobolellus, gen. nov. and Pseudospirobolellus, whereas mtDNA data place this species in the Pachybolidae. Further phylogenetic analyses are needed to explore this apparent incongruence and test the monophyly of Pseudospirobolellidae.

10amitochondrial DNA10anew genus10aphylogeny10aspecies delineation1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://www.publish.csiro.au/is/IS2003102190nas a2200289 4500008004100000022001400041245019200055210006900247260001500316300001000331490000800341520124900349653001001598653001601608653001901624653001301643100002501656700002701681700002401708700002801732700001901760700002001779700001801799700001801817700001801835856004701853 2020 eng d a1313-298900aA new micropolydesmoid millipede of the genus Eutrichodesmus Silvestri, 1910 from Cambodia, with a key to species in mainland Southeast Asia (Diplopoda, Polydesmida, Haplodesmidae)0 anew micropolydesmoid millipede of the genus iEutrichodesmusi Sil c24/11/2020 a59-910 v9963 a

The micropolydesmoid millipede family Haplodesmidae is here recorded from Cambodia for the first time through the discovery of the first, new species of the genus Eutrichodesmus Silvestri, 1910: E. cambodiensis sp. nov. This new species is described from two limestone habitats in Kampot Province, based on abundant material. It is easily distinguished from all related congeners by the following combination of characters: body greyish-brown; limbus roundly lobulate; solenomere partially divided from acropodite by a digitiform lobe, but without hairpad. Brief remarks on the previously-proposed “pecularis-group” are provided and a second group, the “demangei-group”, is established and discussed on the basis of morphological evidence, updating the number of recognised species groups of Eutrichodesmus to two. Detailed morphological illustrations, photographs and a distribution map, as well as remarks on its habitat and mating behaviour of the new species are presented. Furthermore, the current distributions of all 55 presently-known species of Eutrichodesmus are provided and a key to all 23 species that occur in mainland Southeast Asia is given.

10aKarst10anew species10aSoutheast Asia10aTaxonomy1 aSrisonchai, Ruttapon1 aLikhitrakarn, Natdanai1 aSutcharit, Chirasak1 aJeratthitikul, Ekgachai1 aSiriwut, Warut1 aThrach, Phanara1 aChhuoy, Samol1 aNgor, Peng, B1 aPanha, Somsak uhttps://zookeys.pensoft.net/article/57411/02421nas a2200289 4500008004100000022001400041245012500055210006900180260001500249300001200264490000800276520156600284653000901850653001301859653001101872653001401883653000801897100002701905700003001932700001901962700001801981700001801999700002502017700002402042700001802066856004702084 2020 eng d a1313-298900aTwo new species of the millipede genus Plusioglyphiulus Silvestri, 1923 from Cambodia (Diplopoda, Spirostreptida)0 aTwo new species of the millipede genus iPlusioglyphiulusi Silves c04/06/2020 a137-1510 v9383 a

Two new species of Plusioglyphiulus are described from southern Cambodia. Plusioglyphiulus biserratus sp. nov. is clearly distinguished from all congeners by the shape of the telopodites of the posterior gonopods which are distinctly serrate laterally and by the anterior gonopods showing only a pair of single, smooth and curved coxosternal processes. Plusioglyphiulus khmer sp. nov. is distinguished by having most crests on the collum being complete and male legs 1 showing long, prominent, one-segmented telopodites, coupled with the oblong-subtrapeziform, membranous, posterior gonopods with a small bifid process at about a third of the telopodite length. Notes on the variation of Plusioglyphiulus boutini Mauriès, 1970 are also given, including a colour photograph of fresh, live material. A key to all four species of Plusioglyphiulus currently known to occur in Cambodia is also presented.

10aCave10adiplopod10aforest10aIndochina10akey1 aLikhitrakarn, Natdanai1 aGolovatch, Sergei, Ilyich1 aThach, Phanara1 aChhuoy, Samol1 aNgor, Peng, B1 aSrisonchai, Ruttapon1 aSutcharit, Chirasak1 aPanha, Somsak uhttps://zookeys.pensoft.net/article/51234/01781nas a2200265 4500008004100000022001400041245019300055210006900248260001500317300001000332490000800342520087300350653000901223653001301232653003001245653002801275653001301303100002701316700003001343700002801373700002501401700002401426700001801450856004701468 2020 eng d a1313-298900aA remarkable new species of the millipede genus Trachyjulus Peters, 1864 (Diplopoda, Spirostreptida, Cambalopsidae) from Thailand, based both on morphological and molecular evidence0 aremarkable new species of the millipede genus iTrachyjulusi Pete c08/04/2020 a55-720 v9253 a

A new, giant species of Trachyjulus from a cave in southern Thailand is described, illustrated, and compared to morphologically closely related taxa. This new species, T. magnus sp. nov., is much larger than all other congeners and looks especially similar to the grossly sympatric T. unciger Golovatch, Geoffroy, Mauriès & VandenSpiegel, 2012, which is widespread in southern Thailand. Phylogenetic trees, both rooted and unrooted, based on a concatenated dataset of the COI and 28S genes of nine species of Cambalopsidae (Trachyjulus, Glyphiulus, and Plusioglyphiulus), strongly support the monophyly of Trachyjulus and a clear-cut divergence between T. magnus sp. nov. and T. unciger in revealing very high average p-distances of the COI gene (20.80–23.62%).

10aCave10adiplopod10amolecular-based phylogeny10amorphological character10aTaxonomy1 aLikhitrakarn, Natdanai1 aGolovatch, Sergei, Ilyich1 aJeratthitikul, Ekgachai1 aSrisonchai, Ruttapon1 aSutcharit, Chirasak1 aPanha, Somsak uhttps://zookeys.pensoft.net/article/49953/