@article {1520, title = {New taxa of Neotropical Geophilomorpha (Chilopoda)}, journal = {Amazoniana}, volume = {1}, year = {2000}, month = {01/04/2000}, pages = {1-57}, chapter = {1}, abstract = {

A new family (Macronicophilidae) is established for Macronicophilus Silvestri, 1909, currently placed in Geophilidae. Seven new species of Neotropical Geophilomorpha are described: ltyphilus saucius n.sp. and I. sensibilis n.sp. ( Ballophilidae), Hyphydrophilus projectus n.sp. and Ribautia onycophaena n.sp. (Geophilidae), Macronicophilus abbreviatus n.sp., M. unguiseta n.sp. and M. venezolanus n.sp. (Macronicophilidae). The hitherto unknown male of Schendylops marchantariae (Pereira, Minelli \& Barbieri, 1995) is described and two species (Pectiniunguis geayi (Br\ölemann \& Ribaut, 1911) and ltyphilus calinus Chamberlin, 1957 are redescribed from the type and new material. A key to the species of Macronicophilus is provided.

}, keywords = {Amazonia, Chilopoda, Geophilomorpha, Macronicophilidae, Neotropics}, isbn = {0065-6755/2000/001/}, author = {Pereira, Luis Alberto and Foddai, Donatella and Minelli, Alessandro} } @article {1519, title = {First record of Ribautia Br{\"o}lemann, 1909 from Argentina, with description of R. jakulicai sp. n., a new Neotropical member from the Yungas with coxal organs grouped in clusters (Myriapoda: Chilopoda: Geophilomorpha)}, journal = {Studies on Neotropical Fauna and Environment}, volume = {42}, year = {2007}, month = {Aug-01-2007}, pages = {155-168}, chapter = {155}, keywords = {Argentina, Chilopoda, Geophilomorpha, Neotropical Regi{\'o}n, new species, Ribautia, Yungas Forests}, issn = {0165-0521, 1744-5140}, doi = {10.1080/01650520601102377}, url = {http://dx.doi.org/10.1080/01650520601102377}, author = {Pereira, Luis Alberto} } @article {1517, title = {Ribautia lewisi sp. nov., a new centipede from Argentina with unusual tentorial process (Chilopoda: Geophilomorpha, Geophilidae)}, volume = {3630}, year = {2013}, month = {Mar-20-2013}, pages = {225-242}, chapter = {225}, abstract = {

Ribautia lewisi sp. nov. (Chilopoda: Geophilomorpha, Geophilidae) is herein described and illustrated after the holotype (male), paratypes (males and females), and additional non type specimens from Northeastern Argentina (Mesopotamian region). The new species is characterized by having a cluster of coxal organs in each coxopleuron of the ultimate leg-bear-ing segment and a claw-like pretarsus in the ultimate legs, bearing a very unusual feature, in that the internal limbs of ten-torium have a conspicuous tooth-shaped sclerotized process directed inward. R. lewisi sp. nov. is only the second confirmed record of the genus Ribautia from Argentina, the other being R. jakulicai Pereira, 2007 from Northwestern re-gion of the country (Yungas biogeographical province).

}, keywords = {Chilopoda, Geophilidae, Geophilomorpha, Neotropical Regi{\'o}n, new species, Ribautia}, issn = {1175-5334, 1175-5326}, doi = {10.11646/zootaxa.3630.2.2}, url = {http://dx.doi.org/10.11646/zootaxa.3630.2.2}, author = {Pereira, Luis Alberto} } @article {1516, title = {New species and new records of the genus Ribautia Br{\"o}lemann, 1909 (Chilopoda: Geophilomorpha: Geophilidae) from South America}, journal = {Zootaxa}, volume = {1106}, year = {2006}, month = {10-01-2006}, pages = {45}, chapter = {45}, abstract = {

Two new species of Neotropical Geophilomorpha are described: Ribautia combinata n. sp. from Peru and R. donatellae n. sp. from Brazil. New records from Amazonia are given for R. ducalis Pereira, Minelli \& Barbieri, 1995. R. proxima Pereira, Minelli \& Barbieri, 1994 is recorded for the first time from Guyane fran\çaise. Schizoribautia Br\ölemann, 1912 is synonymized under Ribautia Br\ölemann, 1909, syn. nov.

}, keywords = {Chilopoda, Geophilidae, Geophilomorpha, Myriapoda, Neotropical Regi{\'o}n, new records, new species, Ribautia, South America}, issn = {1175-5334, 1175-5326}, doi = {10.11646/zootaxa.1106.1.4}, url = {http://dx.doi.org/10.11646/zootaxa.1106.1.4}, author = {Pereira, Luis A. and Uliana, MARCO and Minelli, Alessandro} } @article {1515, title = {A new species of Ribautia Br{\"o}lemann, 1909 (Chilopoda: Geophilomorpha: Geophilidae) from Peruvian Amazonia, with a key to the Neotropical species of the genus with coxal organs grouped in clusters}, journal = {Studies on Neotropical Fauna and Environment}, volume = {49}, year = {2014}, month = {25/06/2014}, pages = {114-126}, chapter = {114}, abstract = {

Ribautia williamsi sp. nov., a new dwarf geophilomorph centipede from the Lower Urubamba Region, Peruvian
Amazonia, is described and illustrated based on the holotype female. The new species is characterized by having
the coxal organs grouped in clusters (three of these in each coxopleuron of the ultimate leg-bearing segment) and
ventral pore-fields present along all the body; these two combined traits being shared by five other Neotropical
species currently included in the genus Ribautia Br\ölemann, 1909, i.e. R. centralis (Silvestri, 1907) (from Colombia
and Brazil), R. difficilis Pereira, Minelli \& Barbieri, 1995 (from Brazil), R. montana Kraus, 1954 (from Peru), R.
peruana (Verhoeff, 1941) (from Peru), and R. titicacae (Turk, 1955) (from Peru). The new taxon is differentiated
from the aforementioned species by the low number of leg-bearing segments and small body length; it is included
in a key which will enable the identification of all known Neotropical members having coxal organs grouped in
clusters. R. williamsi sp. nov. is the 14th species of Ribautia recorded from Peru.

}, keywords = {Chilopoda, dwarf new species, Geophilidae, Geophilomorpha, Neotropical Regi{\'o}n, Peru, Ribautia}, issn = {0165-0521, 1744-5140}, doi = {https://doi.org/10.1080/01650521.2014.931095}, url = {https://www.tandfonline.com/doi/abs/10.1080/01650521.2014.931095}, author = {Pereira, Luis Alberto} } @article {1403, title = {Ciempi{\'e}s de Sinaloa, M{\'e}xico (Myriapoda: Chilopoda): nuevos registros de especies y ampliaci{\'o}n de su distribuci{\'o}n estatal}, journal = {Dugesiana}, volume = {26}, year = {2019}, month = {25/07/2019}, pages = {79-85}, abstract = {

Resumen: Se han registrado siete especies y una subespecie de ciempi\és en Sinaloa. El estudio de nuevo material, recolectado de 2016 a 2018, permiti\ó identificar 12 especies de ciempi\és, de las cuales ocho representan nuevos registros estatales. Las otras cuatro especies, previamente documentadas para el estado, son registradas en nuevas localidades. Presentamos una lista con los nuevos registros para el estado de Dendrothereua linceci (Wood, 1867), Scolopendra morsitants Linnaeus, 1758, Newportia quadrimeropus (Shelley \& Mercurio, 2005), N. stolli (Pocock, 1896), Pachymerium ferrugineum (Koch, 1835), Mecistocephalus guildingii Newport, 1843, Notiphilides maximiliani (Humbert \& Saussure, 1870) y Orphnaeus brevilabiatus (Newport,1845), as\í como nuevos registros de localidades para Arthrorhabdus pygmaeus (Pocock, 1895), S. polymorpha Wood, 1861, S. viridis Say, 1821 y Rhysida longipes (Newport, 1845). La lista actualizada de especies de ciempi\és para Sinaloa es de 15 especies y una subespecie. En M\éxico, el estado est\á clasificado octavo en riqueza de ciempi\és.

Centipedes of Sinaloa, Mexico (Myriapoda: Chilopoda): new species records and expansion of its regional distribution

Abstract: Seven species and one subspecies of centipedes have been recorded in Sinaloa. The study of new specimens, collected from 2016 to 2018, identified 12 centipedes species, eigth of them are new state records. The other four species, previously recorded in the state, were registered in new localities. We present a list with the new state records of Dendrothereua linceci (Wood, 1867), Scolopendra morsitants Linnaeus, 1758, Newportia quadrimeropus (Shelley \& Mercurio, 2005), N. stolli (Pocock, 1896), Pachymerium ferrugineum (Koch, 1835), Mecistocephalus guildingii Newport, 1843, Notiphilides maximiliani (Humbert \& Saussure, 1870) and Orphnaeus brevilabiatus (Newport,1845), and new localities records from Arthrorhabdus pygmaeus (Pocock, 1895), S. polymorpha Wood, 1861, S. viridis Say, 1821 and Rhysida longipes (Newport, 1845). The current checklist of the Sinaloa state centipedes includes 15 species and one subspecies. In Mexico, the state is ranked eigth in centipedes richness.

}, keywords = {checklist, distribuci{\'o}n, distribution, Geophilomorpha, lista, richness, riqueza, Scolopendromorpha, Scutigeromorpha}, issn = {1405-4094, 2007-9133}, doi = {https://doi.org/10.32870/dugesiana.v26i2.7073}, url = {http://148.202.248.171/dugesiana/index.php/DUG/article/view/7073}, author = {L{\'o}pez-Bonel, Fernando and {\'A}vila-Gaxiola, Claudia Patricia and Cupul-Maga{\~n}a, Fabio G.} } @article {1232, title = {La diversidad de los ciempi{\'e}s (Chilopoda) de M{\'e}xico}, journal = {Dugesiana}, volume = {20}, year = {2013}, month = {30/08/2013}, pages = {17-41}, chapter = {17}, abstract = {

Resumen: Este trabajo presenta un registro hist\órico de la taxonom\ía de los ciempi\és (Chilopoda) descritos para M\éxico. En el pa\ís se han registrado 180 especies (una f\ósil) incluidas en 77 g\éneros, 17 familias y cuatro \órdenes. Veintis\éis g\éneros y 155 especies son end\émicos (una especie f\ósil). S\ólo una familia es end\émica: Eriphantidae (Geophilomorpha). A nivel de g\énero, son reconocibles cuatro elementos biogeogr\áficos en los ciempi\és de M\éxico: end\émico, Hol\ártico, Ne\ártico y Neotropical. Se anexa una lista de las especies de ciempi\és conocidas para el pa\ís, la cual incluye los estados en donde se han recolectado. La diversidad y distribuci\ón de los ciempi\és en M\éxico todav\ía es pobremente conocida y subestimada.

Mexican Diversity of Centipedes (Chilopoda)

Abstract: This paper provides a historical account about of taxonomy of Mexican fauna of centipedes (Chilopoda). A total of 180 identified species (one fossil) in 77 genera, 17 families and four orders have been recorded for Mexico. Twenty-six genera and 155 species are endemic (one fossil). Only one family is endemic: Eriphantidae (Geophilomorpha). At the generic level, four biogeographical elements are recognizable in Mexican centipedes: endemic, Holartic, Neartic, and Neotropical. This work includes a checklist with the centipedes species cited for Mexico and the states of this country where each species has been collected. The diversity and distribution of Mexican centipedes is still poorly known and underestimated.

}, keywords = {Geophilomorpha, Lithobiomorpha, Myriapoda, Scolopendromorpha, Scutigeromorpha, Taxonom{\'\i}a, Taxonomy}, issn = {1405-4094, 2007-9133}, doi = {https://doi.org/10.32870/dugesiana.v20i1.4076}, url = {http://dugesiana.cucba.udg.mx/index.php/DUG/article/view/4076}, author = {Cupul-Maga{\~n}a, Fabio Germ{\'a}n} } @article {1218, title = {Morphology, taxonomy and distribution of Diphyonyx gen. n., a lineage of geophilid centipedes with unusually shaped claws (Chilopoda: Geophilidae)}, journal = {European Journal of Entomology}, volume = {105}, year = {2008}, month = {15/05/2008}, pages = {343-354}, abstract = {

A new genus Diphyonyx is proposed here for a previously unrecognised lineage of geophilid centipedes. It is distinct from other geophilids in its unique combination of morphological characters, including the peculiar shape of the pretarsus of the legs on the anterior part of the trunk. The type species D. conjungens (Verhoeff, 1898), comb. n., is redescribed in detail and its geographical distribution updated (Balkan Peninsula, Anatolia, Crimea). Included in Diphyonyx are also D. sukacevi (Folkmanov\á, 1956), comb. n., and D. garutti (Folkmanov\á \& Dobroruka, 1960), comb. n., both from southern Russia. D. garutti is raised here to species rank.

}, keywords = {Chilopoda, Diphyonyx, Diphyonyx conjungens, geographical distribution, Geophilidae, Geophilomorpha, morphology, new genus, Taxonomy}, issn = {1210-5759}, doi = {10.14411/eje.2008.041}, url = {http://www.eje.cz/artkey/eje-200802-0017_morphology_taxonomy_and_distribution_of_diphyonyx_gen_n_a_lineage_of_geophilid_centipedes_with_unusually_sh.php}, author = {Bonato, Lucio and Zapparoli, Marzio and Minelli, Alessandro} } @article {1027, title = {New data on the family Mecistocephalidae Bollman, 1893 (Chilopoda: Geophilomorpha) from Middle Asia}, journal = {Arthropoda Selecta}, volume = {28}, year = {2019}, abstract = {

The family Mecistocephalidae in Middle Asia contains five accepted species. New records based on material from several collections allow for the distributions of three species to be refined: Arrup asiaticus (Titova, 1975) is new to the fauna of Uzbekistan (Tashkent Region) and to the South Kazakhstan Region of Kazakhstan, A. edentulus (Attems, 1904) is recorded from the Chuy Region of Kyrgyzstan for the first time, while Krateraspis meinerti (Sseliwanoff, 1881) is new to the Jambyl Region of Kazakhstan. Remarks are provided for all of the species encountered, their distributions being mapped as well. A key to all five Mecistocephalidae species occurring in Middle Asia is given.

}, keywords = {Arrup, faunistics, Geophilomorpha, Kazakhstan, Krateraspis, Kyrgyzstan, Mecistocephalidae, new records, Tajikistan., Uzbekistan}, author = {Yurii V. Dyachkov} } @article {958, title = {A new miniature species of geophilomorph centipede from the Ecuadorian Amazon rainforest (Chilopoda: Geophilomorpha: Ballophilidae)}, journal = {Studies on Neotropical Fauna and Environment}, volume = {53}, year = {2018}, month = {Apr-05-2018}, pages = {91 - 106}, abstract = {

A new miniature species of geophilomorph centipede, Ityphilus grismadoi sp. nov. (Chilopoda: Geophilomorpha: Ballophilidae) is fully described and illustrated on the basis of specimens collected in Yasun\í Scientific Station and Jatun Sacha Biological Station, both located in the Ecuadorian Amazon. The new species is characterized by having the internal edge of the forcipular tarsungulum completely smooth, this trait being shared by six other Neotropical members of the genus Ityphilus Cook, 1899; and another species (I. cifuentesi Cupul Maga\ña, 2014) from the transition area between the Neotropical and Nearctic regions. Ityphilus grismadoi sp. nov. is distinguished from these seven species by the low number of leg-bearing segments (37, 39), and the very small body size (7\–9 mm in length) and for a confident identification it is compared in detail with those having a roughly similar range of leg-bearing segments. This is the first record of the genus Ityphilus from Ecuador and the second report of a ballophilid centipede from the country (the other represented by Zygethmus pantenus Chamberlin, 1957). A key to the Neotropical members of Ityphilus with forcipular tarsungulum smooth (including I. cifuentesi) is also given.

}, keywords = {Ballophilidae, Chilopoda, Geophilomorpha, Ityphilus}, issn = {0165-0521}, doi = {10.1080/01650521.2017.1412687}, url = {https://www.tandfonline.com/doi/full/10.1080/01650521.2017.1412687https://www.tandfonline.com/doi/pdf/10.1080/01650521.2017.1412687}, author = {Pereira, Luis Alberto} } @article {957, title = {Una nueva especie de ciempi{\'e}s del g{\'e}nero Ityphilus (Chilopoda: Geophilomorpha: Ballophilidae) de Tamaulipas, M{\'e}xico}, journal = {Revista Mexicana de Biodiversidad}, volume = {85}, year = {2014}, month = {Jan-09-2014}, pages = {723 - 727}, abstract = {

Una nueva especie de ciempi\és del g\énero Ityphilus Cook, 1899, I. cifuentesi sp. nov., se describe a partir de un solo ejemplar macho recolectado en la cueva El Pach\ón, Tamaulipas, M\éxico. Esta es la especie n\úmero 27 descrita para el g\énero a nivel mundial y la segunda para M\éxico. Adem\ás, \éste es el tercer registro de una especie de ciempi\és de la familia Ballophilidae para M\éxico.

}, keywords = {Ballophilidae, Chilopoda, Geophilomorpha, Ityphilus}, issn = {18703453}, doi = {10.7550/rmb.43796}, url = {http://www.revista.ib.unam.mx/index.php/bio/article/view/41}, author = {Cupul-Maga{\~n}a, Fabio Germ{\'a}n} } @article {946, title = {Distribution and indicator value of intertidal centipedes from Mediterranean beaches within and around Port-Cros National Park (Southern France), with proposal of a simplified monitoring (Chilopoda)}, journal = {Bulletin de la Soci{\'e}t{\'e} entomologique de France}, volume = {125}, year = {2020}, month = {20/03/2020}, pages = {41-62}, abstract = {

Abstract: A study on centipedes living on beaches within and around Port-Cros National Park was conducted during the spring of 2019, using hand-collection along transects. One hundred and twenty chilopods (108 of them being Geophilomorpha) were collected, including the halophilous species Henia bicarinata (Dignathodontidae), Geophilus fucorum and Tuoba poseidonis (Geophilidae). The halophilic character of H. bicarinata and G. fucorum was confirmed on the seashore of continental France while T. poseidonis was considered strictly halophilic. Pachymerium ferrugineum (Geophilidae) was considered only halotolerant, in the way it is not restricted to the coastline area. The dominant sediments were observed to have a moderate influence on the presence of halophilous and halobiontic geophilomorphs, with gravel sediments having a more favourable effect. In addition, microhabitats provided by pebbles are of significant importance for halophilic species while the presence of stranded Posidonia seems useful for these species. The availability of driftwood is favourable only when it gives a limited contact with the substrate or the stranded Posidonia layer. Lastly, the impact of human frequentation was very significant regarding halophilic species; all these species were found to be absent where human activity was high. The presence or absence of halophilic centipedes well shows the conservation state of the shore for species living in this habitat. An experimental easy monitoring with identification sheets is given to determine the conservation state of beaches in South-Eastern France.

R\épartition et valeur indicatrice des Chilopodes intertidaux des plages m\éditerran\éennes dans et autour du Parc national de Port-Cros (sud de la France), avec proposition d\’une m\éthode d\’\évaluation simplifi\ée (Chilopoda)

R\ésum\é: Une \étude sur les Chilopodes des plages dans et autour du Parc national de Port-Cros a \ét\é r\éalis\ée au printemps 2019, \à l\’aide de r\écoltes manuelles dans des transects. Cent-vingt chilopodes (108 de ceux-ci \étant des Geophilomorpha) ont \ét\é collect\és, incluant les esp\èces halophiles Henia bicarinata (Dignathodontidae), Geophilus fucorum et Tuoba poseidonis (Geophilidae). Le caract\ère halophile d\’H. bicarinata et de G. fucorum est confirm\é sur le littoral fran\çais, tandis que T. poseidonis est consid\ér\é comme strictement halophile (= halobie). Pachymerium ferrugineum (Geophilidae) est seulement consid\ér\é comme halotol\érant, dans le sens o\ù il n\’est pas restreint aux milieux sal\és littoraux en France. Les s\édiments dominants ont une influence mod\ér\ée sur la pr\ésence des halophiles et halobiontes, avec les s\édiments graveleux ayant un effet plus favorable. En compl\ément, les micro-habitats fournis par les galets sont d\’importance significative pour eux ; quant aux posidonies \échou\ées, elles semblent utiles. La disponibilit\é de bois \échou\és est consid\ér\ée comme favorable uniquement quand ils sont en contact \étroit avec le substrat ou la couche de posidonies \échou\ées. Enfin, l\’impact de la fr\équentation humaine est tr\ès significatif sur les esp\èces halophiles : toutes ces esp\èces sont absentes quand la fr\équentation humaine est \élev\ée. La pr\ésence ou absence des chilopodes halophiles est un bon indicateur de l\’\état de conservation des plages pour les esp\èces d\’Arthropodes inf\éod\ées \à cet habitat. Un suivi exp\érimental simplifi\é avec des fiches d\’identification est donn\é pour d\éterminer l\’\état de conservation des plages dans le sudest de la France.

}, keywords = {Geophilomorpha, halophilic species, Hy{\`e}res Var, Monitoring, wrack}, issn = {0037-928X}, doi = {https://doi.org/10.32475/bsef_2107}, url = {https://www.researchgate.net/publication/340067094_Distribution_and_indicator_value_of_intertidal_centipedes_from_Mediterranean_beaches_within_and_around_Port-Cros_National_Park_Southern_France_with_proposal_of_a_simplified_monitoring_Chilopoda}, author = {Iorio, {\'E}tienne and Geoffroy, David and P{\'e}tillon, Julien} } @article {783, title = {Fine structure and phylogenetic significance of "flexo-canal epidermis glands" in Chilopoda}, journal = {Soil Organisms}, volume = {81}, number = {3}, year = {2009}, pages = {269{\textendash}294}, author = {M{\"u}ller, Carsten H.G. and Rosenberg, J{\"o}rg and Hilken, Gero}, editor = {Xylander, Willi E.R. and Voigtl{\"a}nder, Karin} } @article {768, title = {Tracheal systems in Chilopoda: a comparison under phylogenetic aspects}, journal = {Entomologica Scandinavica Supplement}, volume = {51}, year = {1997}, pages = {49{\textendash}60}, keywords = {Chilopoda, Craterostigmomorpha, Craterostigmus, Cryptopidae, Cryptops, Elektronenmikroskop, G.carpophagus, Geophilidae, Geophilomorpha, H.gabrielis, Himentarium, L.forficatus, Lichtmikroskop, Lithobiidae, Lithobiomorpha, Mikroskopie, morphologie, Myriapoda, Onychophora, Phylogenie, Scolopendra, Scolopendromorpha, Scutigera, Scutigeromorpha, Tracheensystem, vergleichend}, author = {Hilken, Gero} } @article {769, title = {Vergleich von Tracheensystemen unter phylogenetischem Aspekt}, journal = {Verhandlungen des naturwissenschaftlichen Vereins Hamburg (N.F.)}, volume = {37}, year = {1998}, pages = {5{\textendash}94}, keywords = {Chilopoda, Craterostigmomorpha, Craterostigmus, Cryptopidae, Cryptops hortensis, Elektronenmikroskop, G.carpophagus, Geophilidae, Geophilomorpha, H.gabrielis, Himentarium, L.forficatus, Lichtmikroskop, Lithobiidae, Lithobiomorpha, Mikroskopie, morphologie, Phylogenie, S.cingulata, Scolopendra, Scolopendromorpha, Scutigera, Scutigeromorpha, Tracheensystem, vergleichend}, author = {Hilken, Gero} } @article {464, title = {Licht- und elektronenmikroskopische Untersuchungen zur Neurosekretion bei Geophilus longicornis Leach unter besonderer Ber{\"u}cksichtigung der Neuroh{\"a}malorgane}, journal = {Zeitschrift f{\"u}r wissenschaftliche Zoologie (Leipzig)}, volume = {182}, year = {1971}, pages = {62{\textendash}130}, author = {Ernst, Alfred} } @article {449, title = {Morphology of a neglected large-sized species of Geophilus from Iran (Chilopoda: Geophilidae)}, journal = {Zootaxa}, volume = {3736}, year = {2013}, month = {11/10/2013}, pages = {486-500}, abstract = {

Geophilus gigas was described briefly by Attems in 1951 from two sites in Iran, but neither reported nor cited subsequently. The original description is inadequate to understand the full morphology and the taxonomic position of the species. Thus it is here redescribed and figured in detail based on the examination of a syntype and newly collected specimens. Furthermore it is compared with the most similar species of the genus Geophilus. G.\  gigas can be reliably distinguished from other related species for the maximum body size (more than 6 cm), the arrangement and extent of the sternal pore-fields (with additional groups of pores), the coxal pores number and distribution (tens, scattered), and the size of the ulti-mate legs (swollen in both sexes).

}, keywords = {Geophilomorpha, Geophilus gigas, Taxonomy, Western Asia}, issn = {1175-5326}, doi = {10.11646/zootaxa.3736.510.11646/zootaxa.3736.5.4}, url = {https://www.biotaxa.org/Zootaxa/article/view/zootaxa.3736.5.4}, author = {Zarei, Roghaieh and Rahimian, Hassan and Bonato, Lucio} }