01290nas a2200217 4500008004100000022001400041245007400055210006900129260001500198300001000213520065900223653001700882653001500899653000800914653000900922653001300931653000800944100002700952700002000979856007300999 2021 eng d a2118-977300aAtlas of European millipedes 3: Order Chordeumatida (Class Diplopoda)0 aAtlas of European millipedes 3 Order Chordeumatida Class Diplopo c22/09/2021 a1-2443 a

For each of the 534 species of the millipede order Chordeumatida known from Europe, available information on taxonomy, distribution and habitat is summarized, and the distribution in 50 × 50 km UTM/MGRS squares is shown on a map. Comparisons between Chordeumatida and the equally-sized order Julida are made with respect to distribution patterns and history of exploration.

Recommended citation: Kime, R. D. & H. Enghoff (2021): Atlas of European millipedes 3: Order Chordeumatida (Class Diplopoda). European Journal of Taxonomy, 769, 1-244. https://doi.org/10.5852/ejt.2021.769.1497

10adistribution10afaunistics10amap10aMGRS10aTaxonomy10aUTM1 aKime, Richard, Desmond1 aEnghoff, Henrik uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/149701088nas a2200205 4500008004100000022001400041245006700055210006400122260001500186300001000201520049100211653001700702653001500719653000800734653001300742653000800755100002700763700002000790856007200810 2017 eng d a2118-977300aAtlas of European millipedes 2: Order Julida (Class Diplopoda)0 aAtlas of European millipedes 2 Order Julida Class Diplopoda c29/08/2017 a1-2993 a

For each of the 593 species of the millipede order Julida known from Europe, available information on taxonomy, distribution and habitat is summarized, and the distribution in 50 × 50 km UTM squares is shown on a map.

Recommended citation: Kime, R. D. & H. Enghoff (2017): Atlas of European millipedes 2: Order Julida (Class Diplopoda). European Journal of Taxonomy, 346: 1-299. https://doi.org/10.5852/ejt.2017.346

10adistribution10afaunistics10amap10aTaxonomy10aUTM1 aKime, Richard, Desmond1 aEnghoff, Henrik uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/47103170nam a2200157 4500008004100000022003000041245016100071210006900232260003000301300000800331490000600339520257800345100002702923700002002950856004202970 2011 eng d a9789546425782, 954642578800aAtlas of European Millipedes (Class Diplopoda). Volume 1. Orders Polyxenida, Glomerida, Platydesmida, Siphonocryptida, Polyzoniida, Callipodida, Polydesmida0 aAtlas of European Millipedes Class Diplopoda Volume 1 Orders Pol aSofiabPensoft Publishers a2820 v13 a

This is the first in a series of three books which will, for the first time ever, show the distribution of Europe’s more than 1500 species of millipedes on 50 X 50 km square maps. The present volume includes 492 species, including the tiny, beautiful pincushion millipedes, the colourful pill-millipedes and the flat-backed millipedes. The millipede atlas is the result of many years’ meticulous accumulation of records provided by a large number of contributors from all over Europe, interpretation of label data from collections, and scrutiny of old literature. The atlas builds on the taxonomic backbone provided by the Fauna Europaea project which provides information on names, synonyms and distribution by country or region of non-marine European animals.

The decision to produce a series of atlases showing the distribution of myriapods in Europe was taken by the participants at the Fourth International Congress of Myriapodology held in Gargnano in 1978 and soon after this the project received the full backing of the European Invertebrate Survey. Desmond Kime was designated to assemble the data. Much help has been received from the International Centre of Myriapodology in Paris which is acknowledged in this volume, together with the contributions of many members engaged in research in this field. Early in the 21st Century when Henrik Enghoff was playing a major role in Fauna Europaea the two authors joined forces with a view to making this millipede atlas and Fauna Europaea entirely compatible. It is hoped that this atlas will both stimulate and be a foundation for further studies. In several European countries the distribution and often the ecological requirements of millipedes are quite well understood but in many regions the information is fragmentary, the maps revealing where much remains to be done. There are undoubtedly many more species to discover. As a result of the work already undertaken it has become apparent that millipedes are very good bioindicators, being slow-moving and having particular habitat requirements which are helpful in biogeographical and evolutionary studies.

Recommended citation: Kime, R. D. & H. Enghoff (2011): Atlas of European Millipedes (Class Diplopoda). Volume 1. Orders Polyxenida, Glomerida, Platydesmida, Siphonocryptida, Polyzoniida, Callipodida, Polydesmida. Pensoft Publishers, Sofia, 282 pp. ISBN-13: 9789546425782. ISBN-10: 9546425788. https://books.pensoft.net/books/12358

Publication date: 06/2011

 

1 aKime, Richard, Desmond1 aEnghoff, Henrik uhttps://books.pensoft.net/books/1235802524nas a2200229 4500008004100000022002500041245015800066210006900224260001500293300001200308490000900320520176300329653001602092653002302108653002302131653001302154653001302167100002402180700002002204700002302224856004702247 2022 eng d a1313-2970, 1313-298900aTwo new millipede species of the genus Coxobolellus Pimvichai, Enghoff, Panha & Backeljau, 2020 (Diplopoda, Spirobolida, Pseudospirobolellidae)0 aTwo new millipede species of the genus iCoxobolellusi Pimvichai  c09/11/2022 a171-1900 v11283 a

Two new millipede species of the genus Coxobolellus Pimvichai, Enghoff, Panha & Backeljau, 2020 from Thailand are described: Coxobolellus saratani sp. nov. from Loei Province and Coxobolellus serratoligulatus sp. nov. from Uttaradit Province. The descriptions are based on gonopod morphology and two mitochondrial gene fragments (COI and 16S rRNA). The phylogenetic mtDNA analysis assigned the two new species unequivocally to the consistently well-supported Coxobolellus clade, in which they form a fifth subclade that was well supported by maximum likelihood analysis of 16S rRNA, though neither by Bayesian inference nor by COI. The two new Coxobolellus species share four conspicuous gonopodal synapomorphies of the genus: (1) the protruding process on the coxae of the 3rd (and sometimes 4th) pair of male legs, (2) a large, triangular coxae on the 4th–5th pair of legs, (3) a short process of the preanal ring protruding as far as, or slightly beyond, the anal valves, and (4) the posterior gonopod telopodite divided into two parts, with a conspicuous pore opening at the mesal margin at the end of the coxal part of the posterior gonopod. Thus, the two new species provide further evidence of the well-defined monophyly of the genus Coxobolellus. Finally, the paper provides an updated morphological identification key to all currently described Coxobolellus species.

10aDNA barcode10agonopod morphology10aidentification key10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aBackeljau, Thierry uhttps://zookeys.pensoft.net/article/94242/02692nas a2200229 4500008004100000022002500041245014800066210006900214260001500283300001200298490000700310520196600317653000802283653001202291653001402303653001702317100002402334700002002358700001802378700002302396856004302419 2022 eng d a1860-0743, 1435-193500aA new genus of Pseudospirobolellidae (Diplopoda, Spirobolida) from limestone karst areas in Thailand, with descriptions of three new species0 anew genus of Pseudospirobolellidae Diplopoda Spirobolida from li c19/08/2022 a313-3260 v983 a

A new genus of the millipede family Pseudospirobolellidae, Siliquobolellus gen. nov., is described from limestone mountains in Thailand, based on three new species, viz. Siliquobolellus amicusdraconis gen. et sp. nov. from Uthaithani Province, Siliquobolellus constrictus gen. et sp. nov. from Prachuap Khiri Khan Province and Siliquobolellus prasankokae gen. et sp. nov. (type species) from Lampang Province. The descriptions are based on gonopod morphology and mitochondrial DNA data (COI barcodes). The COI barcodes grouped the three new species in a well-supported Siliquobolellus gen. nov. clade. The mean interspecific COI sequence divergence among the three new species was 12% (range: 8–15%). The mean intergeneric COI sequence divergence between Siliquobolellus gen. nov., Coxobolellus Pimvichai, Enghoff, Panha & Backeljau, 2020, and Pseudospirobolellus Carl, 1912 was 19% (range: 14–23%). Three conspicuous gonopodal synapomorphies differentiate Siliquobolellus gen. nov. from other pseudospirobollellid genera: (1) the telopodital part of the posterior gonopod forms a deep concavity, (2) the telopodite of the anterior gonopod is directed distad and does not reach the tip of the coxal part of the anterior gonopod, and (3) the tip of the anterior gonopod coxa is narrowed, curving mesad. As such, the monophyly of the new genus is well supported by both morphological and mitochondrial DNA data. A distribution map and an identification key to the species are provided.

10aCOI10aGonopod10amonophyly10asynapomorphy1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://zse.pensoft.net/article/90032/01599nas a2200217 4500008004100000022002500041245020500066210006900271260001500340300000900355490000900364520084500373653001101218653001201229653001301241653001301254100002401267700002001291700002301311856004701334 2022 eng d a1313-2970, 1313-298900aRedescription of the giant Southeast Asian millipede Spirobolus macrurus Pocock, 1893 and its assignment to the new genus Macrurobolus gen. nov. (Diplopoda, Spirobolida, Pachybolidae)0 aRedescription of the giant Southeast Asian millipede iSpirobolus c22/02/2022 a1-180 v10873 a

A new genus of the millipede family Pachybolidae from Southeast Asia is described: Macrurobolus gen. nov., with Spirobolus macrurus Pocock, 1893 as type species. This latter species is DNA barcoded (COI) and redescribed based on male morphological characters, which hitherto were unknown. The new genus differs from other pachybolid genera by having (1) the preanal ring process long and protruding beyond the anal valves and (2) the anterior gonopod telopodite distally abruptly narrowed, forming an extremely long, slender, elevated process curved caudad. Given that Macrurobolus gen. nov. is a monotypic genus, it is aphyletic and thus requires further taxonomic revision.

10aAphyly10aMyanmar10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aBackeljau, Thierry uhttps://zookeys.pensoft.net/article/71280/03135nas a2200241 4500008004100000022001400041245025000055210006900305260001500374300001200389490000700401520225900408653002202667653001602689653001402705653002002719653002502739100002402764700002002788700001802808700002302826856004402849 2018 eng d a1445-522600aMorphological and mitochondrial DNA data reshuffle the taxonomy of the genera Atopochetus Attems, Litostrophus Chamberlin and Tonkinbolus Verhoeff (Diplopoda: Spirobolida: Pachybolidae), with descriptions of nine new species0 aMorphological and mitochondrial DNA data reshuffle the taxonomy  c01/02/2018 a159-1950 v323 a

Species-level taxonomy and phylogeny of two genera of South-East (SE) Asian pachybolid millipedes are analysed with a combination of morphological characters and DNA sequences (two mitochondrial gene fragments: COI and 16S rRNA). Strong support is found for the genera Litostrophus Chamberlin, 1921 and Atopochetus Attems, 1953 and for a clade consisting of Litostrophus + Atopochetus. Four species of Litostrophus are recognised and (re)described: L. segregatus Chamberlin, 1921 (type species of the genus), L. scaber (Verhoeff, 1938), comb. nov., L. chamaeleon, sp. nov. and L. saraburensis, sp. nov. The genus Tonkinbolus Verhoeff, 1938 (type species T. scaber Verhoeff, 1938) is synonymised under Litostrophus. Atopochetus (type species A. rubropunctatus Attems, 1953), hitherto considered a dubious synonym of Aulacobolus Pocock, 1903, is re-instated for several species until now placed in Tonkinbolus and seven new species. All in all, 10 species of Atopochetus are (re)described: A. dollfusii (Pocock, 1893) (= Aulacobolus rubropunctatus Attems, 1938, syn. nov.), A. moulmeinensis (Pocock, 1893), A. sumatranus (Carl, 1906) (= Trachelomegalus laciniatus Attems, 1937, syn. nov.), A. anaticeps, sp. nov., A. helix, sp. nov., A. setiferus, sp. nov., A. spinimargo, sp. nov., A. truncatus, sp. nov., A. uncinatus, sp. nov. and A. weseneri, sp. nov. Three species recently included in Tonkinbolus are left incertae sedis because they are based exclusively on female type specimens: Spirobolus capucinus Porat, 1896, S. caudulanus Karsch, 1881 and S. macrurus Pocock, 1893. The results of the DNA analysis (COI and 16S rRNA) are congruent with morphological (gonopodal) characters in terms of delimitation of species of Litostrophus and Atopochetus. This is the first DNA study of SE Asian Pachybolidae and as such it provides a basis for further evolutionary and biogeographic studies of SE Asian millipedes.

10amitochondrial DNA10anew species10aphylogeny10aSouth-East Asia10aspecies delimitation1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://www.publish.csiro.au/is/IS1705201621nas a2200241 4500008004100000022001400041245021400055210006900269260001500338300000900353490000800362520077600370653001801146653001501164653001601179653001401195653001301209100002401222700002001246700001801266700002301284856007201307 2016 eng d a2118-977300aA revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part V Nine new spe c24/05/2016 a1-370 v1993 a

The Thyropygus opinatus subgroup (Diplopoda: Harpagophoridae) of the T. allevatus group in Thailand is revised. Based on a phylogenetic analysis of mtDNA sequence data, it is merged with the T. bifurcus subgroup to form an extended T. opinatus subgroup. Nine new species are described: Thyropygus cimi sp. nov. and T. forceps sp. nov. from Nakhonsrithammarat Province, T. culter sp. nov., T. planispina sp. nov., T. undulatus sp. nov. and T. ursus sp. nov. from Krabi Province, T. mesocristatus sp. nov. from Songkhla Province, T. navychula sp. nov. from Phang-Nga Province and T. sutchariti sp. nov. from Phetchaburi Province.

10aDNA barcoding10amillipedes10anew species10aphylogeny10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/32202099nas a2200241 4500008004100000022001400041245011800055210006900173260001500242300001200257490000700269520133500276653001301611653002701624653002201651653001401673653001901687653001301706100002401719700002001743700001801763856007601781 2013 eng d a1095-951300aMolecular phylogeny of the Thyropygus allevatus group of giant millipedes and some closely related groups.0 aMolecular phylogeny of the iThyropygus allevatusi group of giant c20/11/2013 a170-1830 v713 a

Giant cylindrical millipedes of the family Harpagophoridae, especially species of the genus Thyropygus, are broadly distributed in Thailand and nearby countries. They show a great deal of variation in body size, color patterns and gonopodal characters. Phylogenetic analyses of 26 nominate species from six genera in the subfamilies Harpagophorinae and Rhynchoproctinae, as well as nine new morphotypes (regarded as new species), were performed with the DNA sequences from two mitochondrial gene fragments (16S rRNA and COI). The genus Thyropygus (Harpagophorinae) was recovered as monophyletic under all analyses, whilst the representatives of Rhynchoproctinae also formed a monophyletic group. However, the analyses suggested that the tribe Gonoplectini should be elevated to a separate subfamily, Gonoplectinae. The molecular analyses were largely (but not totally) congruent with, and so supported the usefulness of, gonopodal characters for the classification and identification of harpagophorid millipedes, and additionally supported previous studies on the delimitation of species and subgroups. This is the first molecular study inside the family Harpagophoridae and provides the basis for further studies of the evolutionary processes and biogeographic patterns of millipedes in Southeast Asia.

10aendemism10aGeographical isolation10amitochondrial DNA10aphylogeny10aSoutheast Asia10asympatry1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.sciencedirect.com/science/article/abs/pii/S105579031300418101673nas a2200265 4500008004100000022002500041245017100066210006900237260001500306300001000321490000900331520080000340653001301140653001501153653001401168653001601182653001301198100002201211700002401233700002401257700002001281700002701301700001801328856006101346 2012 eng d a1175-5334, 1175-532600aRevision of the rare centipede genus Sterropristes Attems, 1934, with description of a new species from Thailand (Chilopoda: Scolopendromorpha: Scolopendridae)0 aRevision of the rare centipede genus iSterropristesi Attems 1934 c14/09/2012 a35-520 v34843 a

The genus Sterropristes Attems, 1934, currently classified in the scolopendrid subfamily Otostigminae and tribe Sterropristini, is revised. The monotypic genus Malaccolabis Verhoeff, 1937, is synonymized with Sterropristes. Sterropristes sarasinorum Attems, 1934 from Tomohon, Celebes, Indonesia (type species of Sterropristes) and S. metallicus (Verhoeff, 1937) from Penang Hill, Penang, Malaysia (type species of Malaccolabis) are redescribed based on their type material, together with newly collected topotypes of S. metallicus. Sterropristes violaceus Muadsub and Panha n. sp., found under leaves on islands in the Andaman Sea, south-western Thailand, is described.

10aMalaysia10amorphology10aMyriapoda10asystematics10aTaxonomy1 aMuadsub, Sawitree1 aSutcharit, Chirasak1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aEdgecombe, Gregory, D.1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.3484.1.201130nas a2200217 4500008004100000022002500041245014400066210006900210260001500279300001000294490000900304520042000313653001400733653001600747653001300763653001300776100002400789700002000813700001800833856006100851 2011 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 4 the iT cuisi c29/07/2011 a37-480 v29803 a

The Thyropygus cuisinieri subgroup of the T. allevatus group is revised. A new species is described from Thailand: T. jarukchusri n. sp. The following species are re-described: T. cuisinieri Carl, 1917, T. carli Attems, 1938, and T. foliaceus (Demange, 1961), new status (elevated from subspecies status under T. cuisinieri).

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2980.1.301618nas a2200229 4500008004100000022002500041245014300066210006900209260001500278300001000293490000900303520087700312653001401189653001601203653002001219653001301239653001301252100002401265700002001289700001801309856006101327 2011 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 3 the iT indur c06/07/2011 a47-680 v29413 a

The Thyropygus induratus subgroup of the T. allevatus group is revised. Three new species are described from Thailand: T. dormiens n. sp., T. laterolobatus n. sp. and T. macrosiamensis n. sp. The following species are redescribed and re-elevated from subspecies status under T. allevatus (Karsch, 1881): Thyropygus induratus Attems, 1936 (= T. punctatus Attems, 1938, new synonym), Thyropygus quietus Attems, 1938, Thyropygus resimus Attems, 1938, and Thyropygus uncinatus (Demange, 1961). Thyropygus siamensis Verhoeff, 1938 (= Thyropisthus ligulus Demange, 1961, new synonym), is re-described. A case of species with identical gonopods but very different body size (T. induratus and T. quietus) is discussed.

10amillipede10anew species10asize difference10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2941.1.301583nas a2200169 4500008004100000022001400041245023000055210006900285260001500354300001000369490000700379520092100386100002401307700002001331700001801351856004401369 2010 eng d a1445-522600aThe Rhynchoproctinae, a south-east Asiatic subfamily of giant millipedes: cladistic analysis, classification, four new genera and a deviating new species from north-west Thailand (Diplopoda : Spirostreptida : Harpagophoridae)0 aRhynchoproctinae a southeast Asiatic subfamily of giant milliped c17/05/2010 a51-800 v243 a

The subfamily Rhynchoproctinae is revised at the generic level. Four new genera and a remarkable new species are described: Alienostreptus, gen. nov. from Vietnam, Armatostreptus, gen. nov., Heptischius, gen. nov., and Heptischius lactuca, sp. nov. from Thailand, and Prominulostreptus, gen. nov. from China. Agaricogonopus Zhang & Zhang, 1997, is removed from synonymy with Junceustreptus Demange, 1961. All 14 genera of the subfamily share three diagnostic gonopodal characters: the posterior surface of the anterior coxal fold forms a deep concavity, there is a mesal flap on the gonopod coxa, and the posterior coxal fold is very low. A cladistic analysis based on morphological data supports monophyly of Rhynchoproctinae and the distincness of the newly described genera. A dichotomous key to the genera is included.

1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.publish.csiro.au/is/is0905201206nas a2200217 4500008004100000022002500041245014200066210006900208260001500277300000900292490000900301520049900310653001400809653001600823653001300839653001300852100002400865700002000889700001800909856006100927 2009 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 2: the T. bifurcus subgroup (Diplopoda, Spirostreptida, Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 2 the iT bifur c22/07/2009 a1-150 v21653 a

The Thyropygus bifurcus subgroup of the T. allevatus group is revised. Four new species are described from Thailand: T. demangei n. sp., T. quadricuspis n. sp., T. richardhoffmani n. sp., from Trang province and T. casjeekeli n. sp., from Krabi province. The other species of the T. bifurcus subgroup, viz., T. bifurcus (Demange, 1986), and T. enghoffi (Demange, 1989), are redescribed.

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2165.1.101898nas a2200217 4500008004100000022002500041245014200066210006900208260001500277300001000292490000900302520119000311653001401501653001601515653001301531653001301544100002401557700002001581700001801601856006101619 2009 eng d a1175-5334, 1175-532600aA revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)0 arevision of the iThyropygus allevatusi group Part 1 the iT opina c20/02/2009 a17-500 v20163 a

The Thyropygus opinatus subgroup of the T. allevatus group is revised. The T. opinatus subgroup corresponds to the genus Cornugonus Demange, 1961, which is formally synonymized under Thyropygus. Eight new species are described from Thailand: T. bearti n. sp., T. brachyacanthus n. sp., T. loxia n. sp., from Suratthani province, T. bispinus n. sp., from Uthaithani and Phrae provinces, T. bispinispatula n. sp., from Chumphon province, T. chelatus n. sp., from Nakhonsrithammarat province, T. cristagalli n. sp., from Phang Nga province and T. erectus n. sp., from Satun province. The other species of the T. opinatus subgroup, viz., T. opinatus (Karsch, 1881), n. comb., T. floweri (Demange, 1961), n. comb., T. implicatus (Demange, 1961), n. comb., and T. inflexus (Demange, 1989), n. comb., are redescribed. New records are given for T. opinatus, T. implicatus and T. floweri. The type locality of T. floweri is shown to be in Thailand rather than Malaysia.

10amillipede10anew species10aTaxonomy10aThailand1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak uhttps://www.mapress.com/zt/article/view/zootaxa.2016.1.201510nas a2200193 4500008004100000022002500041245005600066210005400122300001100176490000800187520095400195653001501149653001301164653001201177100002001189700002501209700002001234856006201254 2016 eng d a0012-8317, 0012-831700aChecklist of the millipedes (Diplopoda) of Tanzania0 aChecklist of the millipedes Diplopoda of Tanzania a51-1130 v1053 a

A checklist of millipedes (Diplopoda) known to occur in Tanzania is given. Based on all available literature and abundant hitherto unpublished material, 296 species of millipedes are recorded, including seven species never before recorded from the country, viz., Helicochetus digititarsus Kraus, 1957, Helicochetus gregorii (Pocock, 1896), Geotypodon intermedius (Carl, 1909), Plethocrossus nairobinus Attems, 1914, Lophostreptus bicolor Carl, 1909, Oreiadessa diana Hoffman, 1990, and Chondromorpha xanthotricha (Attems, 1898). Eight species erroneously recorded from Tanzania are excluded from the list. A few records from Kenya and Uganda are also given, including the first record of Otostreptus gilvitarsus (Attems, 1914) from Kenya. A historical account of millipede collecting in Tanzania is included.

 

10acollecting10aendemism10aSPECIES1 aEnghoff, Henrik1 aHoffman, Richard, L.1 aHowell, Kim, M. uhttps://www.ajol.info/index.php/jeanh/article/view/13967104609nas a2200241 4500008004100000022002500041245014900066210006900215260001500284300000900299490000800308520382800316653002704144653001104171653002604182653001204208653001404220653002704234653001304261100002004274700002004294856005304314 2011 eng d a1313-2970, 1313-298900aOn some surface structures of potential taxonomic importance in families of the suborders Polydesmidea and Dalodesmidea (Polydesmida, Diplopoda)0 asome surface structures of potential taxonomic importance in fam c20/12/2011 a1-240 v1563 a

Surface structures have rarely been the subject of a comprehensive study in Polydesmida despite their tremendous variety within this order. A number of these peripheral structures are here studied in most families of the suborders Polydesmidea and Dalodesmidea (sensu Hoffman 1980), using scanning electron microscopy. An illustrated description of the surface sculpture of the prozonite, the limbus and the intercalary cuticular micro-scutes on the metazonite is given for the first time for the studied families, together with an account of some other poorly known surface structures. Taken together, these characters allow us to recognize two main groupings of families. The families Ammodesmidae, Cryptodesmidae, Cyrtodesmidae, Haplodesmidae, Oniscodesmidae and Pyrgodesmidae have knobs on the posterior part of the prozonites, a toothed to lobed limbus, and no micro-scutes on the metazonites, wheras the families Fuhrmannodesmidae, Polydesmidae, Dalodesmidae, Macrosternodesmidae, Nearctodesmidae, Opisotretidae and Trichopolydesmidae have no knobs on the posterior part of the prozonites, a spiky or reduced limbus, and intercalary micro-scutes on the metazonites. The results are complemented with literature records and compared with current taxonomic and phylogenetic interpretations of the group.

 

10acuticular micro-scutes10alimbus10ametazonital outgrowth10aozopore10aprozonite10asurface microsculpture10aTaxonomy1 aAkkari, Nesrine1 aEnghoff, Henrik uhttps://zookeys.pensoft.net/articles.php?id=225901309nas a2200217 4500008004100000022002500041245009400066210006900160260001500229300001100244490000800255520066300263653001500926653001100941653001000952653001300962653001600975100002600991700002001017856005401037 2018 eng d a1313-2970, 1313-298900aFirst continental troglobiont Cylindroiulus millipede (Diplopoda, Julida, Julidae)0 aFirst continental troglobiont iCylindroiulusi millipede Diplopod c08/11/2018 a93-1030 v7953 a

The new species of millipede Cylindroiulus villumi is described from a cave in the Estremenho karst massif in central Portugal. It is the first cave-adapted species of its genus with a strict subterranean life-style in continental Europe, and is the fifth blind species of the genus. The new species is illustrated with photographs and diagrammatic drawings. It is tentatively placed in the purely Iberian Cylindroiulus perforatus-group. The differences between the new species and its relatives are discussed, as well as its adaptations to a subterranean life-style.

10aCave fauna10aJulida10aKarst10aPortugal10atroglobiont1 aReboleira, Ana, SP.S.1 aEnghoff, Henrik uhttps://zookeys.pensoft.net/articles.php?id=2761901533nas a2200193 4500008004100000022002500041245014700066210006900213260001500282300001200297490000900309520088200318653001701200653001301217653001201230100002001242700002001262856005701282 2008 eng d a1175-5334, 1175-532600aCylindroiulus mitta n.sp., a new Tunisian millipede (Diplopoda, Julida: Julidae) and notes on the Cylindroiulus distinctus group0 aiCylindroiulus mittai nsp a new Tunisian millipede Diplopoda Jul c14/05/2008 a61–680 v17683 a

Abstract: Cylindroiulus mitta n.sp. is described from Cap Bon, northeastern Tunisia, and its assignment to the North African C. distinctus group is discussed. Descriptions of the female vulvae of other group members are presented, as well as illustrations of gonopods and vulvae of further Cylindroiulus species from Tunisia.

Résumé: Cylindroiulus mitta n. sp. est décrit de la région du Cap Bon, nord-est de Tunisie et son appartenance au groupe Nord africain C. distinctus est discutée. Des descriptions des vulves de femelles des autres espèces du groupe sont ici présentées ainsi que des illustrations des gonopodes et des vulves d’autres espèces du genre Cylindroiulus de Tunisie.

10aNorth Africa10aTaxonomy10aTunisia1 aAkkari, Nesrine1 aEnghoff, Henrik uhttps://mapress.com/zt/article/view/zootaxa.1768.1.401997nas a2200169 4500008004100000245010300041210006900144260001500213300001000228490000700238520143800245100002901683700001701712700002301729700002001752856005501772 2021 eng d00aPenetrative and non-penetrative interaction between Laboulbeniales fungi and their arthropod hosts0 aPenetrative and nonpenetrative interaction between Laboulbeniale c12/11/2021 a1–70 v113 a

Laboulbeniales are a highly specialized group of fungi living only on arthropods. They have high host specificity and spend their entire life-cycle on an arthropod host. Taxonomic characters of Laboulbeniales are based on the architecture of the cells of the parenchymal thallus, i.e. the visible part of the fungus outside the host. The extent of the fungus spreading inside the host—the haustorium—remains largely unknown. The attachment to the arthropod host is fundamental to understand the fungus-animal interaction, but how this truly occurs is unclear. Recent evidences question the strictly parasitic life-style of Laboulbeniales. We used micro-computed tomography (µCT) and 3D reconstructions to visualize, for the first time, the complete structure of Laboulbeniales species in situ on their hosts. We compared the haustoriate species, Arthrorhynchus nycteribiae on an insect host to the non-haustoriate species, Rickia gigas on a millipede host. Our results confirm that some Laboulbeniales species are ectoparasitic and have a haustorial structure that penetrates the host’s cuticle, while others are ectobionts and are only firmly attached to the host’s cuticle without penetrating it. The presence and the morphology of the haustorium are important traits for Laboulbeniales evolution, and key factors for future understanding of host dependence and specificity.

1 aReboleira, Ana, Sofia PS1 aMoritz, Leif1 aSantamaría, Sergi1 aEnghoff, Henrik uhttps://www.nature.com/articles/s41598-021-01729-x01990nas a2200241 4500008004100000022001400041245012300055210006900178260001500247300001200262490000800274520123800282653001701520653001201537653001401549653001801563653001501581100002201596700002001618700001701638700002001655856007301675 2021 eng d a2118-977300aIntegrative description of new giant pill-millipedes from southern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)0 aIntegrative description of new giant pillmillipedes from souther c11/08/2021 a108-1320 v7623 a

Two new species of giant pill-millipedes, Zephronia viridisoma Rosenmejer & Wesener sp. nov. and Sphaerobelum aesculus Rosenmejer & Wesener sp. nov., are described based on museum samples from southern Thailand. Zephronia viridisoma sp. nov. comes from Khao Lak, while the type locality of S. aesculus sp. nov. is on Phuket Island. Both species are described integratively, combining light microscopy, scanning electron microscopy, multi-layer photography, micro-CT scans and genetic barcoding. Genetic barcoding was successfully conducted for holotypes of both new species, which could be added to a dataset of all published sequences of the family Zephroniidae, including all described species from Thailand, Laos and Cambodia up to 2020. Genetic barcoding of the COI gene revealed another female of S. aesculus sp. nov., 160 km east of the type locality. Both new species are genetically distant from all other Zephroniidae from Thailand and surrounding countries, showing uncorrected p-distances of 16.8–23.1%. A virtual cybertype of a paratype of Z. viridisoma sp. nov. was created and made publically accessible.

10abiodiversity10aCT scan10acybertype10aDNA barcoding10aSoil fauna1 aRosenmejer, Trine1 aEnghoff, Henrik1 aMoritz, Leif1 aWesener, Thomas uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/145700785nas a2200169 4500008004100000245018800041210006900229260001200298100002900310700001800339700003300357700003000390700002000420700002000440700002500460856013000485 2012 eng d00aDaily Activities of the Giant Pill-Millipede Zephronia cf. viridescens Attems, 1936 (Diplopoda: Sphaerotheriida: Zephroniidae) in a Deciduous Forest in Northern Thailand0 aDaily Activities of the Giant PillMillipede iZephroniai cf iviri c01/20121 aWongthamwanich, Nattarin1 aPanha, Somsak1 aSitthicharoenchai, Duangkhae1 aPradatsundarasar, Art-Ong1 aSeelanan, Tosak1 aEnghoff, Henrik1 aThirakhupt, Kumthorn uhttps://myriatrix.myspecies.info/content/daily-activities-giant-pill-millipede-zephronia-cf-viridescens-attems-1936-diplopoda01066nas a2200193 4500008004100000022001400041245015500055210006900210260001500279300001200294490000900306520040200315653001800717653001600735653001300751100002200764700002000786856006600806 2021 eng d a1175-532600aA new giant species of the millipede genus Prionopetalum Attems, 1909 from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)0 anew giant species of the millipede genus iPrionopetalumi Attems  c31/03/2021 a561-5700 v49503 a

A new large species of the millipede genus Prionopetalum Attems, 1909, P. nessiae sp. nov. from Mt. Kilimanjaro, Tanzania, is described, with SEM imaging and illustrations of relevant taxonomic characters, and mapped. Comments on the significance of sternum 9 in descriptions of Odontopygidae, and on the intrageneric differences in Prionopetalum, are included.

10anew character10anew species10aTaxonomy1 aRosenmejer, Trine1 aEnghoff, Henrik uhttps://www.biotaxa.org/Zootaxa/article/view/zootaxa.4950.3.801978nas a2200229 4500008004100000022001400041245017600055210006900231260001500300300001000315490000700325520122700332653001701559653001401576653001501590653001401605653001301619100002001632700002001652700002901672856004701701 2008 eng d a1445-522600aPachybolini – a tribe of giant Afrotropical millipedes: arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae)0 aPachybolini a tribe of giant Afrotropical millipedes arguments f c18/03/2008 a37-530 v223 a

Madabolus, gen. nov. is described to accommodate M. maximus, sp. nov. (type species) from several localities in western Madagascar. A phylogenetic analysis of 26 spirobolidan taxa, based on morphological characters, places Madabolus, gen. nov. with the Afrotropical genera Pelmatojulus de Saussure, 1860; Pachybolus Cook, 1897; Hadrobolus Cook, 1897 and Epibolus Cook, 1897. The tribe Pachybolini is proposed to include Madabolus, gen. nov. and the four abovementioned genera. Putative autapomorphies of Pachybolini are: a closed incisura lateralis; setae of gnathochilarial lamella lingualis apical; vulva kidney- or crescent-shaped, with valves meeting in a sinuous fissure and crest protruding. Within Pachybolini, Pelmatojulus (West Africa) stands out by virtue of an increased number of apical antennal sensory cones, with great probability an apomorphy. Madabolus, gen. nov. and east African genera of the tribe share another potential synapomorphy, namely, an increased number of setae on the gnathochilarial lamellae linguales. Other Malagasy genera of Pachybolidae do not seem to be related to Afrotropical genera.

10abiogeography10aMadabolus10amorphology10aphylogeny10aTaxonomy1 aWesener, Thomas1 aEnghoff, Henrik1 aWägele, Johann-Wolfgang uhttp://www.publish.csiro.au/?paper=IS0700801672nas a2200205 4500008004100000022001400041245013300055210006900188260001500257300000900272520098600281653002601267653000801293653001601301653001301317100002101330700002201351700002001373856007301393 2020 eng d a2118-977300aA mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)0 amountain of millipedes IX Species of the family Gomphodesmidae f c29/06/2020 a1-353 a

A new genus and six new species of the family Gomphodesmidae from the Udzungwa Mts are described, including Pogoro alopias Rosenmejer & Enghoff sp. nov., Pogoro siren Rosenmejer & Enghoff sp. nov., Pogoropsis prolixopes Rosenmejer & Enghoff gen. et sp. nov., Emphysemastix frampt Olsen & Enghoff sp. nov., Agrophogonus hamulus Olsen & Enghoff sp. nov., and Agrophogonus pusillokiellandi Olsen & Enghoff sp. nov. Emphysemastix dracarys Olsen & Enghoff sp. nov. from Iringa city is described. Descriptive notes are given for Pogoro scharffi Hoffman, 2005, and Agrophogonus mwanihana Hoffmann, 2005. General gomphodesmid gonopod morphology is described and illustrated. A key to Udzungwa gomphodesmids is presented, as well as revised keys to all species of Emphysemastix and Agrophogonus. All gomphodesmid species from the Udzungwa Mts are mapped.

10aEastern Arc Mountains10akey10anew species10aTaxonomy1 aOlsen, Sissel, A1 aRosenmejer, Trine1 aEnghoff, Henrik uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/100501911nas a2200181 4500008004100000022001400041245025200055210006900307260001500376300000900391520116900400653001601569653002301585653001601608653001301624100002001637856007201657 2020 eng d a2118-977300aA mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited – a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts and introduction of the first pair of male legs as a source of taxonomic characters0 amountain of millipedes VIII The genus iAquattuori Frederiksen 20 c14/04/2020 a1-323 a

Full title: A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited – a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)

A large, newly collected material of the genus Aquattuor from the Udzungwa Mts, Tanzania, has been studied. Two new species are described, viz., A. mollilobus sp. nov. from the Udzungwa Mts and A. nguruensis sp. nov. from the Nguru Mts, Tanzania. Aquattuor claudiahempae Enghoff & Frederiksen, 2015, is recorded as new for the Udzungwa fauna. Characters from the first pair of male legs are illustrated for all Aquattuor species and are shown to be useful for species discrimination. A new gonopodal difference between A. submajor Enghoff, 2015 and A. udzungwensis Enghoff, 2015 is described and illustrated. Intrageneric relationships and distribution patterns are discussed. A distribution map and a key to all Aquattuor species are presented.

10aEastern Arc10aidentification key10anew species10aTaxonomy1 aEnghoff, Henrik uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/90101661nas a2200229 4500008004100000022001400041245012600055210006900181260001500250300000900265490000800274520092200282653001601204653001901220653001301239100002401252700002001276700002001296700002201316700002001338856007301358 2020 eng d a2118-977300aA strikingly coloured new giant millipede from Vietnam has copycat in Borneo (Diplopoda, Spirostreptida, Harpagophoridae)0 astrikingly coloured new giant millipede from Vietnam has copycat c20/10/2020 a1-150 v7223 a

A new giant species of the subfamily Rhynchoproctinae with strikingly bi-coloured, red-yellow legs contrasting with a black body is described: Alienostreptus bicoloripes sp. nov. from Vietnam. The new species is assigned to the genus Alienostreptus Pimvichai, Enghoff & Panha, 2010 hitherto comprising one species, A. alienus (Attems, 1936), and differing from other rhynchoproctine genera by having the femoral spine duplicated. Species of this genus share three synapomorphic characters of the subfamily, viz (1) anterior coxal fold forms deep concavity in posterior view, (2) posterior coxal fold very low, and (3) posterior coxal fold with mesal flap. A superficially very similar colourful species from Borneo, also with bi-coloured legs but clearly not belonging to Alienostreptus due to the position of ventral soft pads on male legs, is documented based on photographs.

10anew species10aSoutheast Asia10aTaxonomy1 aPimvichai, Piyatida1 aAkkari, Nesrine1 aSemenyuk, Irina1 aConstant, Jerôme1 aEnghoff, Henrik uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/112901947nas a2200241 4500008004100000022001400041245016500055210006900220260001500289300001200304490000700316520117700323653001301500653001401513653001201527653001001539653001601549100002201565700002001587700002701607700002401634856004701658 2020 eng d a1445-522600aPhylogenetic placement of the monotypic genus Holopodostreptus Carl, 1913 and notes on the systematics of Pseudonannolenidae (Spirostreptida: Cambalidea)0 aPhylogenetic placement of the monotypic genus iHolopodostreptusi c14/08/2020 a661-6770 v343 a

The genus Holopodostreptus Carl, 1913 is a poorly known monotypic genus endemic to Ecuador. It differs from most other Pseudonannolenidae by a set of characteristics of the gnathochilarium and gonopods. Herein, we provide descriptive notes of males of H. braueri Carl, 1913 based on recently collected material. Females of the species are described for the first time, and images of the vulvae are provided. In addition, we attempt for the first time to place Holopodostreptus in a phylogenetic context. With a total of 14 taxa and 41 morphological characters, and under implied weighting (k = 6), a single most-parsimonious tree is recovered (total fit = 35.86). The genus is recovered as sister-group of Physiostreptus Silvestri, 1903. The clade Holopodostreptus + Physiostreptus is supported by one synapomorphy (absence of setae on the prefemoral process of leg-pair 1 in males) and one homoplastic transformation (narrow telopodite). Based on the phylogenetic results, notes on the subfamilies Pseudonannoleninae, Cambalomminae and Physiostreptinae and a key for all pseudonannolenid genera are provided.

10aColombia10aDiplopoda10aEcuador10aHaiti10aWest Indies1 aIniesta, Luiz, FM1 aEnghoff, Henrik1 aBrescovit, Antonio, D.1 aBouzan, Rodrigo, S. uhttp://www.publish.csiro.au/?paper=IS2001202849nas a2200229 4500008004100000022001400041245017100055210006900226260001500295300001200310490000700322520208700329653002202416653001402438653001402452653002402466100002402490700002002514700001802534700002302552856004402575 2020 eng d a1445-522600aIntegrative taxonomy of the new millipede genus Coxobolellus, gen. nov. (Diplopoda : Spirobolida : Pseudospirobolellidae), with descriptions of ten new species0 aIntegrative taxonomy of the new millipede genus iCoxobolellusi g c14/08/2020 a591-6170 v343 a

Pseudospirobolellidae is a poorly known family of spirobolidan millipedes with only two genera and five described species. Yet, the descriptive taxonomy and molecular systematics of this group have been largely neglected. Therefore, the present work presents an integrative taxonomic study of new pseudospirobolellid taxa in Thailand. To this end, two mitochondrial gene fragments (COI and 16S rRNA) combined with morphological characters were used to define the genus Coxobolellus, gen. nov. with 10 new species, viz. C. albiceps, sp. nov., C. compactogonus, sp. nov., C. fuscus, sp. nov., C. nodosus, sp. nov., C. serratus, sp. nov., C. simplex, sp. nov., C. tenebris, sp. nov., C. tigris, sp. nov., C. transversalis, sp. nov. and C. valvatus, sp. nov. The interspecific COI sequence divergences among the new species ranged from 6 to 15%. The intergeneric COI sequence divergence between species of Coxobolellus, gen. nov., Benoitolus birgitae and Pseudospirobolellus sp. ranged from 20 to 23%. Three major morphological differences separate Coxobolellus, gen. nov. from Benoitolus and Pseudospirobolellus, namely (1) the protruding process on the 3rd (and 4th) coxae on male legs, (2) the posterior gonopod telopodite divided into two parts, and (3) a conspicuous opening pore at the mesal margin at the end of the coxal part of the posterior gonopod. Thus, the new genus is well supported by both mtDNA and morphological evidence, while the delimitation of the 10 new species is supported by the congruence between mtDNA and morphological data. Yet, with respect to the relationships of Benoitolus birgitae, morphological data suggest a similarity with Coxobolellus, gen. nov. and Pseudospirobolellus, whereas mtDNA data place this species in the Pachybolidae. Further phylogenetic analyses are needed to explore this apparent incongruence and test the monophyly of Pseudospirobolellidae.

10amitochondrial DNA10anew genus10aphylogeny10aspecies delineation1 aPimvichai, Piyatida1 aEnghoff, Henrik1 aPanha, Somsak1 aBackeljau, Thierry uhttps://www.publish.csiro.au/is/IS2003101012nas a2200193 4500008004100000022001400041245017700055210006900232260001500301300000900316520032300325653001000648653001700658653001300675653001600688100002000704700002200724856007200746 2020 eng d a2118-977300aThe first blind spirostreptid millipede, found in a cave in Morocco; with notes on the genus Odontostreptus Attems, 1914 (Diplopoda, Spirostreptida, Spirostreptidae)0 afirst blind spirostreptid millipede found in a cave in Morocco w c17/06/2020 a1-113 a

The first eyeless species of the suborder Spirostreptidea, Odontostreptus fadriquei sp. nov., is described from a cave in Morocco. The new species, which exhibits some troglomorphic traits, is compared with the highly variable O. maroccanus (Attems, 1914) of which new material is presented.

10acaves10aNorth Africa10aTaxonomy10atroglobiont1 aEnghoff, Henrik1 aReboleira, Ana, S uhttps://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/98502116nas a2200241 4500008004100000022001400041245020600055210006900261260001500330300001200345490000900357520125800366653001401624653002201638653001901660653001601679653002201695653002201717100002301739700002601762700002001788856006601808 2020 eng d a1175-532600aDescription of the female morphology of the relict troglobiont millipede Cantabrodesmus lorioli Mauriès 1971 (Diplopoda, Polydesmida, Chelodesmidae), and new records in the deepest pit of Spain0 aDescription of the female morphology of the relict troglobiont m c16/12/2020 a421-4300 v48953 a

The troglobiont millipede, Cantabrodesmus lorioli Mauriès, 1971, inhabits a few caves in a small region of Northern Spain. It is considered a relict species because of its geographical isolation compared to others in its family, and its to some extent uncertain systematic position. Cantabrodesmus lorioli has been placed in three different families, and now is considered a member of the Chelodesmidae, a family with only one other known native European representative. There are few records of this species, and almost nothing is known about its biology. The only available information on this species is the description of the male and a list of caves where it has been captured. In the present work, we provide the first description of the female morphology, illustrated by scanning electron microscopy, and a series of new records from caves in a region where it had never been captured before, including the deepest known pit in Spain, Torca del Porrón. The female morphology does not show many differences from that of the male, but several characters are discussed. All of the known information on the morphology of the species is used to discuss the implications for its biology and systematic position.

10aDiplopoda10afemale morphology10ahypogean fauna10anew records10aTorca del Porrón10atroglobiomorphism1 aGilgado, José, D.1 aAguinaga, José, A. N1 aEnghoff, Henrik uhttps://www.biotaxa.org/Zootaxa/article/view/zootaxa.4895.3.700495nas a2200133 4500008004100000245010700041210006900148260000900217490000700226100002000233700002000253700002400273856006400297 2014 eng d00aSegmentation of the millipede trunk as suggested by a homeotic mutant with six extra pairs of gonopods0 aSegmentation of the millipede trunk as suggested by a homeotic m c20140 v111 aAkkari, Nesrine1 aEnghoff, Henrik1 aMinelli, Alessandro uhttp://www.biomedcentral.com/content/pdf/1742-9994-11-6.pdf00636nas a2200169 4500008004100000020001400041245016900055210006900224260001400293300001300307490000700320100002000327700002000347700002400367700002300391856005200414 2015 en d a1932-620300aA New Dimension in Documenting New Species: High-Detail Imaging for Myriapod Taxonomy and First 3D Cybertype of a New Millipede Species (Diplopoda, Julida, Julidae)0 aNew Dimension in Documenting New Species HighDetail Imaging for  c2015-8-26 ae01352430 v101 aAkkari, Nesrine1 aEnghoff, Henrik1 aMetscher, Brian, D.1 aEvans, Alistair, R uhttp://dx.plos.org/10.1371/journal.pone.013524300729nas a2200169 4500008004100000020002500041245026000066210006900326260001500395300000900410490000800419100002000427700002300447700002000470700001700490856005200507 2013 eng d a1313-2970, 1313-298900aRevolving SEM images visualising 3D taxonomic characters: application to six species of the millipede genus Ommatoiulus Latzel, 1884, with description of seven new species and an interactive key to the Tunisian members of the genus (Diplopoda, Juli0 aRevolving SEM images visualising 3D taxonomic characters applica c2013-09-03 a5-450 v3281 aAkkari, Nesrine1 aCheung, David, K-B1 aEnghoff, Henrik1 aStoev, Pavel uhttp://zookeys.pensoft.net/articles.php?id=361004944nas a2200241 4500008004100000022001400041245014100055210006900196260001500265300001200280490000700292520419400299653001704493653001304510653001504523653001504538653002104553653001604574100002004590700002004610700002004630856005204650 2009 eng d a1313-298900aReview of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of 'fire millipedes' (Diplopoda)0 aReview of the Spirobolida on Madagascar with descriptions of twe c04/09/2009 a1 - 1280 v193 a

Twelve new genera and 37 new species of Spirobolida are described: Corallobolus cruentus gen. n., sp. n., Sanguinobolus maculosus gen. n., sp. n., Colossobolus semicyclus gen. n., sp. n., C. oblongopedus sp. n., C. giganteus sp. n., C. minor sp. n., C. litoralis sp. n., C. aculeatus sp. n., C. pseudoaculeatus sp. n., Zehntnerobolus gen. n., Flagellobolus pauliani gen. n., sp. n., Riotintobolus mandenensis gen. n., sp. n., R. minutus sp. n., R. aridus sp. n., R. anomalus sp. n., Pseudocentrobolus aureus gen. n., sp. n., P. vohibasiensis sp. n., Granitobolus endemicus gen. n., sp. n., G. andohahelensis sp. n., Caprobolus andringitra gen. n., sp. n., Alluviobolus laticlavius gen. n., sp. n., A. tsimelahy sp. n., A. antanosy sp. n., Ostinobolus rufus gen. n., sp. n., O. stellaris sp. n., O. montanus sp. n., O. subterraneus sp. n., and Hylekobolus brachiosauroides gen. n., sp. n., H. rufus sp. n., H. griseus sp. n., H. albicollaris sp. n., H. goodmani sp. n., H. montanus sp. n., H. analavelona sp. n., H. latifrons sp. n., H. andasibensis sp. n., H. marojejy sp. n., H. anjanaharibe sp. n. All genera and species are endemic to Madagascar. Hylekobolus belongs to the family Spirobolellidae, while all other Malagasy genera of Spirobolida belong to the Pachybolidae. Among them, only Zehntnerobolus gen. n. is based on a previously described species: Spirobolus rubripes de Saussure & Zehntner, 1897, whereas the remaining 11 new genera altogether contain (a total of) 37 new species. Three of the new genera are large-bodied “fire millipedes” (>100 mm long) with striking red/black colour patterns. The new discoveries increase the number of endemic Malagasy genera of Spirobolida more than fivefold (from 3 to 15). The number of endemic species recorded from Madagascar has more than doubled (to 61). Body length of the new species varies greatly (between 23 and 170 mm). Keys to all Malagasy Spirobolida families, genera, as well as the newly described species, are provided. Four species, Rhinocricus collaris de Saussure & Zehntner, 1897, Spirobolus grandidieri de Saussure & Zehntner, 1897, Spirobolus olympiaceus Karsch, 1881, and Spirobolus sikorae de Saussure & Zehntner, 1897 are only known from females and are listed as nomina dubia. Some genera occur in the dry or spiny forest as well as in the rainforest ecosystem; species, however, are restricted to a single habitat type. In the rainforest, up to six different genera were collected in the same individual forest. Numerous genera are adapted to a special ecological niche. A high proportion of species are microendemic, often to some tiny, highly endangered forests. Of special conservation concern are the species which occur only in Analavelona, Ambatotsirongorongo, the subhumid, globally unique littoral forest of Petriky and the littoral rainforests of Mandena and Sainte Luce.

10abiogeography10aendemism10aMadagascar10amillipedes10aniche separation10aSpirobolida1 aWesener, Thomas1 aEnghoff, Henrik1 aSierwald, Petra uhttp://zookeys.pensoft.net/articles.php?id=203701630nas a2200229 4500008004100000022001400041245009000055210006900145260001500214300001400229490000800243520089700251653001501148653001001163653002201173653001001195653002001205100002301225700002001248700002601268856010601294 2014 eng d a0027-551400aLaboulbeniales on millipedes: the genera Diplopodomyces and Troglomyces0 aLaboulbeniales on millipedes the genera iDiplopodomycesi and iTr c19/09/2014 a1027-10380 v1063 a

The genera Diplopodomyces and Troglomyces (Laboulbeniales, Ascomycota) parasitizing millipedes (Diplopoda) are reviewed. We describe four new species: D. lusitanipodos parasitic on Lusitanipus from Portugal and on Cyphocallipus from Spain, D. veneris on Lusitanipus from Portugal, T. botryandrus on Leptoiulus from Italy and T. rossii on Ophyiulus, Leptoiulus and Typhloiulus from Italy. Diplopodomyces veneris is a dimorphic species, with different morphotypes growing on the male copulatory organs and on the surroundings of the female gonopore respectively. The type species of both genera also have been studied, their descriptions are revised and a neotype for T. manfrediae is designated. New records from Italy are given for D. callipodos and T. manfrediae.

10aAscomycota10acaves10aIberian Peninsula10aItaly10aparasitic fungi1 aSantamaría, Sergi1 aEnghoff, Henrik1 aReboleira, Ana, SP.S. uhttps://www.tandfonline.com/doi/full/10.3852/13-381https://www.tandfonline.com/doi/pdf/10.3852/13-38102073nas a2200205 4500008004100000022001400041245021500055210007100270260001500341300001200356490000700368520133400375653001601709653001501725653001301740653001601753100002001769700002301789856005501812 2015 eng d a1439-609200aInfectious intimacy and contaminated caves—three new species of ectoparasitic fungi (Ascomycota: Laboulbeniales) from blaniulid millipedes (Diplopoda: Julida) and inferences about their transmittal mechanisms0 aInfectious intimacy and contaminated caves—three new species of  c22/03/2015 a249-2630 v153 a

Laboulbeniales is an order of more than 2000 species of small ascomycete fungi which are ectoparasites of insects, millipedes and mites. They are often highly host-specific and often are also highly specific with regard to which body parts they infect. Laboulbeniales from millipedes are particularly poorly known—only 12 species have been described until now, mostly from Europe. Here, we describe the first laboulbeniaceous parasites from blaniulid millipedes: Troglomyces bilabiatus from Acipes spp., Troglomyces pusillus from Iberoiulus cavernicola Ceuca, 1967 and Troglomyces triandrus from Archiboreoiulus palidus (Brade-Birks, 1920). The generic description of Troglomyces is emended. Different types of site specificity of the three new parasites, as well as of previously described species, are explained in terms of aspects of host biology: mating behaviour and habitat. Species from epigean hosts tend to show a high degree of site specificity suggesting transfer during host copulation, whereas most species from cave hosts show no pronounced site specificity and are probably transferred via the substrate. Possible roles of host defensive secretions and parthenogenesis in relation to infection with Laboulbeniales are briefly discussed.

10aBlaniulidae10aParasitism10aTaxonomy10aTroglomyces1 aEnghoff, Henrik1 aSantamaría, Sergi uhttp://link.springer.com/10.1007/s13127-015-0208-801934nas a2200241 4500008004100000022001400041245011500055210007100170260001500241300001200256490000800268520119100276653001501467653001401482653001101496653002001507653001601527653001901543100002301562700002001585700002201605856006501627 2016 eng d a1179-315500aHidden biodiversity revealed by collections-based research—Laboulbeniales in millipedes: genus Rickia0 aHidden biodiversity revealed by collectionsbased research—Laboul c12/01/2016 a101-1270 v2433 a

Nine new species of the genus Rickia parasitizing Diplopoda are described: R. appendicifera from Iran on Chiraziulus, R. candelabriformis from Australia and New Zealand on several Spirostreptida, R. galatheae from New Zealand on Eumastigonus, R. gigas from Tanzania on ‘Spirostreptus’ and Archispirostreptus, R. lophophora from Tanzania on some genera of Spirostreptidae, R. obelostrepti from Tanzania on Obelostreptus, R. odontopygiidarum from Tanzania on Odontopygidae, R. platessa from Singapore and Thailand on Pachybolidae, and R. rhynchophora from Australia on Trigoniulus.  This nearly triples the previously known number of species of Rickia species on millipedes; all 14 species are treated in this study with photomicrographs, additional observations on previously described taxa and comments. A key to these species is also provided. The Laboulbeniales species were found on millipede specimens preserved mostly at the Natural History Museum of Denmark, from samples collected between 1916 to the present.

10aAscomycota10aDiplopoda10aJulida10aparasitic fungi10aSpirobolida10aSpirostreptida1 aSantamaría, Sergi1 aEnghoff, Henrik1 aReboleira, Ana, S uhttps://biotaxa.org/Phytotaxa/article/view/phytotaxa.243.2.102202nas a2200157 4500008004100000245014000041210006900181260001500250300000900265490000700274520164000281100002701921700002001948700002301968856005301991 2018 eng d00aNovelty upon novelty visualized by rotational scanning electron micrographs (rSEM): Laboulbeniales on the millipede order Chordeumatida0 aNovelty upon novelty visualized by rotational scanning electron  c28/11/2018 a1-160 v133 a

Laboulbeniales are highly specific ectoparasitic fungi of arthropods (insects, millipedes, and arachnids). The first Laboulbeniales parasitizing the millipede order Chordeumatida (Diplopoda) were discovered and described as a new dioecious genus of Laboulbeniales, Thaxterimyces, to accommodate the new species T. baliensis. Also the millipede host is a new species and is described as Metopidiothrix sheari. This is the first time Laboulbeniales fungus and its millipede host are described as new together. Males of Metopidiothrix have the most extensive secondary sexual modifications in the entire class Diplopoda. Although nothing is known about the function of these modifications, the unique pattern of Laboulbeniales infection in the new millipede species is obviously related to host sexual behavior. Rotational Scanning Electron Micrographs (rSEM) are used to create a 3D comprehensive model to examine the fungal-host interaction, a more advanced visualization of the ectoparasitic fungus on its host. Laboulbeniales diversity on millipedes is still understudied, and a consistent effort is needed to unveil and understand the extent and diversity of this biological interaction. Due to their minute size and apparently non-detrimental effect on their hosts, Laboulbeniales in general have been largely ignored by mycologists and neglected by generations of entomologists. As a result a significant component of global biodiversity has been strongly underestimated, and a wealth of new discoveries is still to be made both in the field and in existing museum collections.

1 aReboleira, Ana, SP. S.1 aEnghoff, Henrik1 aSantamaría, Sergi uhttps://dx.plos.org/10.1371/journal.pone.020690002266nas a2200217 4500008004100000022001400041245012100055210006900176260001500245300000900260520154200269653001901811653002401830653001901854653001501873653001601888100002301904700002001927700002601947856007501973 2018 eng d a2118-977300aNew species of Troglomyces and Diplopodomyces (Laboulbeniales, Ascomycota) from millipedes (Diplopoda)0 aNew species of iTroglomycesi and iDiplopodomycesi Laboulbeniales c27/04/2018 a1-203 a

We describe five new species of fungi of the order Laboulbeniales Lindau growing on millipedes and belonging to the genera Diplopodomyces W.Rossi & Balazuc and Troglomyces S.Colla. Three new species of Diplopodomyces, viz. Diplopodomyces coronatus Santam., Enghoff & Reboleira sp. nov. living on Serboiulus spelaeophilus Gulicka, 1967 from Bulgarian caves, Diplopodomyces liguliphorus Santam., Enghoff & Reboleira sp. nov. on an unidentified species of Spirobolida from Sri Lanka, and Diplopodomyces ramosus Santam., Enghoff & Reboleira sp. nov. on Pachyiulus spp. from Turkey, Macedonia and Serbia; and two new species of Troglomyces, viz. Troglomyces dioicus Santam., Enghoff & Reboleira sp. nov. on Nepalmatoiulus sp. from Myanmar, and Troglomyces tetralabiatus Santam., Enghoff & Reboleira sp. nov. on Caucaseuma Strasser, 1970 and Heterocaucaseuma Antić & Makarov, 2016 from caves in Western Caucasus. Troglomyces dioicus sp. nov. is the first dioecious species described in the genus Troglomyces. Keys for all hitherto known species of Diplopodomyces and Troglomyces are presented, as is a discussion of the status of both genera. Additional records for Diplopodomyces lusitanipodos Santam., Enghoff & Reboleira and Troglomyces manfrediae S.Colla are also included.

10aDiplopodomyces10aidentification keys10aLaboulbeniales10amillipedes10aTroglomyces1 aSantamaría, Sergi1 aEnghoff, Henrik1 aReboleira, Ana, SP.S. uhttp://www.europeanjournaloftaxonomy.eu/index.php/ejt/article/view/55801442nas a2200229 4500008004100000022001400041245012200055210006900177260001500246300001000261490000700271520070900278653003000987653003101017653001401048653002001062653001701082100002301099700002001122700002201142856004801164 2020 eng d a1314-405700aThe first Laboulbeniales (Ascomycota, Laboulbeniomycetes) from an American millipede, discovered through social media0 afirst Laboulbeniales Ascomycota Laboulbeniomycetes from an Ameri c14/05/2020 a45-530 v673 a

Laboulbeniales are highly specialized arthropod-associated fungi. The majority of the almost 2200 known species live on insects, although they also occur on other arthropod hosts. Recently, the number of Laboulbeniales associated with millipedes has increased considerably. Here we describe the first species of a Laboulbeniales fungus, Troglomyces twitteri sp. nov., from an American millipede. The new species was initially discovered on a photo of Cambala annulata (Say, 1821) from Ohio, USA, which had been shared on Twitter. A subsequent microscopic study of Cambala millipedes in museum collections in Denmark and France confirmed the discovery.

10aanimal-fungus interaction10acollections-based research10aDiplopoda10aLaboulbeniaceae10asocial media1 aSantamaría, Sergi1 aEnghoff, Henrik1 aReboleira, Ana, S uhttps://mycokeys.pensoft.net/article/51811/01609nas a2200217 4500008004100000022001400041245026200055210007000317260001500387300001200402490000900414520074500423653001801168653002401186653003801210653001601248100002301264700002001287700002501307856005901332 2015 eng d a1175-532600aThe hypogean Iberian genus Typhlopsychrosoma Mauriès, 1982 (Diplopoda, Chordeumatida, Vandeleumatidae): distribution map, key to species, first record in a Mesovoid Shallow Substratum (MSS) and detailed iconography of T. baeticaense (Mauriès, 2013)0 ahypogean Iberian genus Typhlopsychrosoma Mauriès 1982 Diplopoda  c25/03/2015 a337-3460 v39373 a

The troglobiont millipede Typhlopsychrosoma baeticaense (Mauriès, 2013) is recorded in the Mesovoid Shallow Substratum (MSS) of two screes in the Aitana and Bernia mountains (Eastern Iberian Peninsula), far away from its known distribution area. A detailed Scanning Electron Microscopy iconography provides additional information on gonopod morphology, as well as other details such as the inconspicuous evaginations of the cuticle at the place of the eyes in the anophthalmous specimens. We present an updated distribution map and a key to species of the genus, with illustrations of the gonopods of all species described so far. The implications of the appearance of this hypogean species in an MSS are discussed.

10aBaetic System10ahypogean millipedes10asuperficial subterranean habitats10atroglobiont1 aGilgado, José, D.1 aEnghoff, Henrik1 aOrtuño, Vicente, M. uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.3937.2.502189nas a2200253 4500008004100000022001400041245018800055210006900243260001600312300001200328490000800340520127600348653002301624653002501647653002401672653001801696653002301714653003801737100002301775700002001798700002001818700002501838856007201863 2015 eng d a0044-523100aHidden biodiversity in the Iberian Mesovoid Shallow Substratum (MSS): New and poorly known species of the millipede genus Archipolydesmus Attems, 1898 (Diplopoda, Polydesmidae)0 aHidden biodiversity in the Iberian Mesovoid Shallow Substratum M cJan-09-2015 a13 - 380 v2583 a

Systematic sampling of the Mesovoid Shallow Substratum (MSS) at several sites in three regions of the Iberian Peninsula has revealed two new and one very poorly known species of the genus Archipolydesmus Attems, 1898. These are the first records of Archipolydesmus in the MSS, although the genus had previously been recorded from several caves in southern Spain. Archipolydesmus foliatus sp. nov. shows inter-population variability between different mountain massifs of the province of Alicante, as well as intra-population variability at some sites; A. altibaeticus sp. nov. exclusively inhabits the MSS of the highest screes of Sierra Nevada (3000 masl) in the province of Granada, and it closely resembles Archipolydesmus maroccanus Attems, 1898; from North Africa; Archipolydesmus bedeli (Brölemann, 1902 )had only been re-collected once since its description, and the male, hitherto unknown, is described here.Detailed SEM documentation of the main characters of each species is provided, and a discussion on theimplications of these findings in the MSS is provided. Also, a distribution map, an updated key to species of the genus in English, and a brief review of millipedes found in the MSS are provided.

10aAlicante mountains10aGuadarrama mountains10ahypogean millipedes10aSierra Nevada10asubterranean fauna10asuperficial subterranean habitats1 aGilgado, José, D.1 aEnghoff, Henrik1 aTinaut, Alberto1 aOrtuño, Vicente, M. uhttps://www.sciencedirect.com/science/article/pii/S004452311500057102017nas a2200241 4500008004100000022001400041245025200055210006900307260001600376300001200392490000900404520110200413653002401515653000801539653002101547653003801568100002301606700002001629700002001649700002401669700002501693856005701718 2015 eng d a1175-532600aSierra Nevada (Granada, Spain): a high-altitude biogeographical crossroads for millipedes (Diplopoda), with first data on its MSS fauna and description of a new species of the genus Ceratosphys Ribaut, 1920 (Chordeumatida: Opisthocheiridae)0 aSierra Nevada Granada Spain a highaltitude biogeographical cross cJul-11-2016 a391-4100 v40443 a

Millipedes (Diplopoda), with a few notable exceptions, are poor dispersers, showing a very high degree of endemicity, not the least in mountains. The first samplings of the Mesovoid Shallow Substratum (MSS) of the higher altitudes of the Sierra Nevada Mountains (Baetic System, Southern Spain) have led to the discovery of a high number of millipedes, each of the species present showing a different degree of establishment in this subterranean environment. An update of the knowledge on the millipedes of this region, the first data of the millipede communities in the MSS and the description of Ceratosphys cryodeserti Gilgado, Mauriès & Enghoff n. sp. are here provided, as well as the first data on the humidity and temperature fluctuations in the MSS of this high mountain. The new species is similar to other Baetico-Riffan species, while the only previously known congener from the region, C. soutadei Mauriès, 1969, has more similarities to certain Pyrenean species. Biogeographical relationships of all the captured species are also discussed.

10ahypogean millipedes10aMSS10aOrophilous fauna10asuperficial subterranean habitats1 aGilgado, José, D.1 aEnghoff, Henrik1 aTinaut, Alberto1 aMauriès, Jean-Paul1 aOrtuño, Vicente, M. uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.4044.301870nas a2200253 4500008004100000022001400041245014800055210006900203260001600272300001200288490000900300520103300309653001501342653001601357653001301373653002501386653003001411100002301441700002101464700002001485700002401505700002501529856006201554 2017 eng d a1175-532600aA new genus and species of Haplobainosomatidae (Diplopoda: Chordeumatida) from the MSS of the Sierra de Guadarrama National Park, central Spain0 anew genus and species of Haplobainosomatidae Diplopoda Chordeuma cFeb-11-2018 a492-5100 v43473 a

The chordeumatidan fauna of the Iberian Peninsula is far from being well known, but recent efforts are improving that knowledge. Samplings carried out in the Milieu Souterrain Superficiel (also known as the Mesovoid Shallow Substratum) on several screes of the Sierra de Guadarrama National Park (central Spain) have led to the discovery of a new species of chordeumatidan millipede. This new species belongs to the family Haplobainosomatidae, which is mainly known from the northern Iberian Peninsula, and thus this is the first record of the family in central Spain. However, the extremely simple gonopods of this species are so different from all other known species of the family that it must be placed in a new genus. In this work, a complete description of the species, named as Guadarramasoma ramosae gen. & sp. nov., with a detailed iconography based on scanning electron microscopy images is provided together with a distribution map and a brief discussion of the implications of this new finding.

10amillipedes10anew species10aorobiome10aSierra de Guadarrama10aSubterranean environments1 aGilgado, José, D.1 aLedesma, Enrique1 aEnghoff, Henrik1 aMauriès, Jean-Paul1 aOrtuño, Vicente, M. uhttps://biotaxa.org/Zootaxa/article/view/zootaxa.4347.3.401405nas a2200241 4500008004100000022001400041245016500055210006900220260001600289300001200305490000900317520050100326653001600827653001900843653001000862653001300872653001900885100002000904700002300924700002500947700002000972856017100992 2018 eng d a1175-532600aOut of the dark void: Ommatoiulus longicornis n. sp., a new julid from Spain (Diplopoda, Julida) with notes on some troglobiomorphic traits in millipedes0 aOut of the dark void iOmmatoiulus longicornisi n sp a new julid  cMay-05-2019 a415-4290 v44203 a

A new millipede species, Ommatoiulus longicornis n. sp. is described from the subterranean habitat in eastern Spain, and the elongation of its antennae is discussed in relation to the habitat where it dwells. We take the occasion to discuss the elongation of appendages as a troglobiomorphic character in millipedes and shed the light on the various, sometimes controversial, hypotheses that could have led to these modifications in millipedes and other subterranean arthropods.

10anew species10aSchizophyllini10aSpain10aTaxonomy10atroglomorphism1 aAkkari, Nesrine1 aGilgado, José, D.1 aOrtuño, Vicente, M.1 aEnghoff, Henrik uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4420.3https://biotaxa.org/Zootaxa/article/view/zootaxa.4420.3.7https://biotaxa.org/Zootaxa/article/viewFile/38811/33160