02116nas a2200241 4500008004100000022001400041245020600055210006900261260001500330300001200345490000900357520125800366653001401624653002201638653001901660653001601679653002201695653002201717100002301739700002601762700002001788856006601808 2020 eng d a1175-532600aDescription of the female morphology of the relict troglobiont millipede Cantabrodesmus lorioli Mauriès 1971 (Diplopoda, Polydesmida, Chelodesmidae), and new records in the deepest pit of Spain0 aDescription of the female morphology of the relict troglobiont m c16/12/2020 a421-4300 v48953 a

The troglobiont millipede, Cantabrodesmus lorioli Mauriès, 1971, inhabits a few caves in a small region of Northern Spain. It is considered a relict species because of its geographical isolation compared to others in its family, and its to some extent uncertain systematic position. Cantabrodesmus lorioli has been placed in three different families, and now is considered a member of the Chelodesmidae, a family with only one other known native European representative. There are few records of this species, and almost nothing is known about its biology. The only available information on this species is the description of the male and a list of caves where it has been captured. In the present work, we provide the first description of the female morphology, illustrated by scanning electron microscopy, and a series of new records from caves in a region where it had never been captured before, including the deepest known pit in Spain, Torca del Porrón. The female morphology does not show many differences from that of the male, but several characters are discussed. All of the known information on the morphology of the species is used to discuss the implications for its biology and systematic position.

10aDiplopoda10afemale morphology10ahypogean fauna10anew records10aTorca del Porrón10atroglobiomorphism1 aGilgado, José, D.1 aAguinaga, José, A. N1 aEnghoff, Henrik uhttps://www.biotaxa.org/Zootaxa/article/view/zootaxa.4895.3.701609nas a2200217 4500008004100000022001400041245026200055210007000317260001500387300001200402490000900414520074500423653001801168653002401186653003801210653001601248100002301264700002001287700002501307856005901332 2015 eng d a1175-532600aThe hypogean Iberian genus Typhlopsychrosoma Mauriès, 1982 (Diplopoda, Chordeumatida, Vandeleumatidae): distribution map, key to species, first record in a Mesovoid Shallow Substratum (MSS) and detailed iconography of T. baeticaense (Mauriès, 2013)0 ahypogean Iberian genus Typhlopsychrosoma Mauriès 1982 Diplopoda  c25/03/2015 a337-3460 v39373 a

The troglobiont millipede Typhlopsychrosoma baeticaense (Mauriès, 2013) is recorded in the Mesovoid Shallow Substratum (MSS) of two screes in the Aitana and Bernia mountains (Eastern Iberian Peninsula), far away from its known distribution area. A detailed Scanning Electron Microscopy iconography provides additional information on gonopod morphology, as well as other details such as the inconspicuous evaginations of the cuticle at the place of the eyes in the anophthalmous specimens. We present an updated distribution map and a key to species of the genus, with illustrations of the gonopods of all species described so far. The implications of the appearance of this hypogean species in an MSS are discussed.

10aBaetic System10ahypogean millipedes10asuperficial subterranean habitats10atroglobiont1 aGilgado, José, D.1 aEnghoff, Henrik1 aOrtuño, Vicente, M. uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.3937.2.502189nas a2200253 4500008004100000022001400041245018800055210006900243260001600312300001200328490000800340520127600348653002301624653002501647653002401672653001801696653002301714653003801737100002301775700002001798700002001818700002501838856007201863 2015 eng d a0044-523100aHidden biodiversity in the Iberian Mesovoid Shallow Substratum (MSS): New and poorly known species of the millipede genus Archipolydesmus Attems, 1898 (Diplopoda, Polydesmidae)0 aHidden biodiversity in the Iberian Mesovoid Shallow Substratum M cJan-09-2015 a13 - 380 v2583 a

Systematic sampling of the Mesovoid Shallow Substratum (MSS) at several sites in three regions of the Iberian Peninsula has revealed two new and one very poorly known species of the genus Archipolydesmus Attems, 1898. These are the first records of Archipolydesmus in the MSS, although the genus had previously been recorded from several caves in southern Spain. Archipolydesmus foliatus sp. nov. shows inter-population variability between different mountain massifs of the province of Alicante, as well as intra-population variability at some sites; A. altibaeticus sp. nov. exclusively inhabits the MSS of the highest screes of Sierra Nevada (3000 masl) in the province of Granada, and it closely resembles Archipolydesmus maroccanus Attems, 1898; from North Africa; Archipolydesmus bedeli (Brölemann, 1902 )had only been re-collected once since its description, and the male, hitherto unknown, is described here.Detailed SEM documentation of the main characters of each species is provided, and a discussion on theimplications of these findings in the MSS is provided. Also, a distribution map, an updated key to species of the genus in English, and a brief review of millipedes found in the MSS are provided.

10aAlicante mountains10aGuadarrama mountains10ahypogean millipedes10aSierra Nevada10asubterranean fauna10asuperficial subterranean habitats1 aGilgado, José, D.1 aEnghoff, Henrik1 aTinaut, Alberto1 aOrtuño, Vicente, M. uhttps://www.sciencedirect.com/science/article/pii/S004452311500057102017nas a2200241 4500008004100000022001400041245025200055210006900307260001600376300001200392490000900404520110200413653002401515653000801539653002101547653003801568100002301606700002001629700002001649700002401669700002501693856005701718 2015 eng d a1175-532600aSierra Nevada (Granada, Spain): a high-altitude biogeographical crossroads for millipedes (Diplopoda), with first data on its MSS fauna and description of a new species of the genus Ceratosphys Ribaut, 1920 (Chordeumatida: Opisthocheiridae)0 aSierra Nevada Granada Spain a highaltitude biogeographical cross cJul-11-2016 a391-4100 v40443 a

Millipedes (Diplopoda), with a few notable exceptions, are poor dispersers, showing a very high degree of endemicity, not the least in mountains. The first samplings of the Mesovoid Shallow Substratum (MSS) of the higher altitudes of the Sierra Nevada Mountains (Baetic System, Southern Spain) have led to the discovery of a high number of millipedes, each of the species present showing a different degree of establishment in this subterranean environment. An update of the knowledge on the millipedes of this region, the first data of the millipede communities in the MSS and the description of Ceratosphys cryodeserti Gilgado, Mauriès & Enghoff n. sp. are here provided, as well as the first data on the humidity and temperature fluctuations in the MSS of this high mountain. The new species is similar to other Baetico-Riffan species, while the only previously known congener from the region, C. soutadei Mauriès, 1969, has more similarities to certain Pyrenean species. Biogeographical relationships of all the captured species are also discussed.

10ahypogean millipedes10aMSS10aOrophilous fauna10asuperficial subterranean habitats1 aGilgado, José, D.1 aEnghoff, Henrik1 aTinaut, Alberto1 aMauriès, Jean-Paul1 aOrtuño, Vicente, M. uhttp://biotaxa.org/Zootaxa/issue/view/zootaxa.4044.301870nas a2200253 4500008004100000022001400041245014800055210006900203260001600272300001200288490000900300520103300309653001501342653001601357653001301373653002501386653003001411100002301441700002101464700002001485700002401505700002501529856006201554 2017 eng d a1175-532600aA new genus and species of Haplobainosomatidae (Diplopoda: Chordeumatida) from the MSS of the Sierra de Guadarrama National Park, central Spain0 anew genus and species of Haplobainosomatidae Diplopoda Chordeuma cFeb-11-2018 a492-5100 v43473 a

The chordeumatidan fauna of the Iberian Peninsula is far from being well known, but recent efforts are improving that knowledge. Samplings carried out in the Milieu Souterrain Superficiel (also known as the Mesovoid Shallow Substratum) on several screes of the Sierra de Guadarrama National Park (central Spain) have led to the discovery of a new species of chordeumatidan millipede. This new species belongs to the family Haplobainosomatidae, which is mainly known from the northern Iberian Peninsula, and thus this is the first record of the family in central Spain. However, the extremely simple gonopods of this species are so different from all other known species of the family that it must be placed in a new genus. In this work, a complete description of the species, named as Guadarramasoma ramosae gen. & sp. nov., with a detailed iconography based on scanning electron microscopy images is provided together with a distribution map and a brief discussion of the implications of this new finding.

10amillipedes10anew species10aorobiome10aSierra de Guadarrama10aSubterranean environments1 aGilgado, José, D.1 aLedesma, Enrique1 aEnghoff, Henrik1 aMauriès, Jean-Paul1 aOrtuño, Vicente, M. uhttps://biotaxa.org/Zootaxa/article/view/zootaxa.4347.3.401405nas a2200241 4500008004100000022001400041245016500055210006900220260001600289300001200305490000900317520050100326653001600827653001900843653001000862653001300872653001900885100002000904700002300924700002500947700002000972856017100992 2018 eng d a1175-532600aOut of the dark void: Ommatoiulus longicornis n. sp., a new julid from Spain (Diplopoda, Julida) with notes on some troglobiomorphic traits in millipedes0 aOut of the dark void iOmmatoiulus longicornisi n sp a new julid  cMay-05-2019 a415-4290 v44203 a

A new millipede species, Ommatoiulus longicornis n. sp. is described from the subterranean habitat in eastern Spain, and the elongation of its antennae is discussed in relation to the habitat where it dwells. We take the occasion to discuss the elongation of appendages as a troglobiomorphic character in millipedes and shed the light on the various, sometimes controversial, hypotheses that could have led to these modifications in millipedes and other subterranean arthropods.

10anew species10aSchizophyllini10aSpain10aTaxonomy10atroglomorphism1 aAkkari, Nesrine1 aGilgado, José, D.1 aOrtuño, Vicente, M.1 aEnghoff, Henrik uhttps://biotaxa.org/Zootaxa/issue/view/zootaxa.4420.3https://biotaxa.org/Zootaxa/article/view/zootaxa.4420.3.7https://biotaxa.org/Zootaxa/article/viewFile/38811/33160