@article {1782, title = {Morphological and mitochondrial DNA data reshuffle the taxonomy of the genera Atopochetus Attems, Litostrophus Chamberlin and Tonkinbolus Verhoeff (Diplopoda: Spirobolida: Pachybolidae), with descriptions of nine new species}, journal = {Invertebrate Systematics}, volume = {32}, year = {2018}, month = {01/02/2018}, pages = {159-195}, type = {Research Article}, abstract = {

Species-level taxonomy and phylogeny of two genera of South-East (SE) Asian pachybolid millipedes are analysed with a combination of morphological characters and DNA sequences (two mitochondrial gene fragments: COI and 16S rRNA). Strong support is found for the genera Litostrophus Chamberlin, 1921 and Atopochetus Attems, 1953 and for a clade consisting of Litostrophus + Atopochetus. Four species of Litostrophus are recognised and (re)described: L. segregatus Chamberlin, 1921 (type species of the genus), L. scaber (Verhoeff, 1938), comb. nov., L. chamaeleon, sp. nov. and L. saraburensis, sp. nov. The genus Tonkinbolus Verhoeff, 1938 (type species T. scaber Verhoeff, 1938) is synonymised under Litostrophus. Atopochetus (type species A. rubropunctatus Attems, 1953), hitherto considered a dubious synonym of Aulacobolus Pocock, 1903, is re-instated for several species until now placed in Tonkinbolus and seven new species. All in all, 10 species of Atopochetus are (re)described: A. dollfusii (Pocock, 1893) (= Aulacobolus rubropunctatus Attems, 1938, syn. nov.), A. moulmeinensis (Pocock, 1893), A. sumatranus (Carl, 1906) (= Trachelomegalus laciniatus Attems, 1937, syn. nov.), A. anaticeps, sp. nov., A. helix, sp. nov., A. setiferus, sp. nov., A. spinimargo, sp. nov., A. truncatus, sp. nov., A. uncinatus, sp. nov. and A. weseneri, sp. nov. Three species recently included in Tonkinbolus are left incertae sedis because they are based exclusively on female type specimens: Spirobolus capucinus Porat, 1896, S. caudulanus Karsch, 1881 and S. macrurus Pocock, 1893. The results of the DNA analysis (COI and 16S rRNA) are congruent with morphological (gonopodal) characters in terms of delimitation of species of Litostrophus and Atopochetus. This is the first DNA study of SE Asian Pachybolidae and as such it provides a basis for further evolutionary and biogeographic studies of SE Asian millipedes.

}, keywords = {mitochondrial DNA, new species, phylogeny, South-East Asia, species delimitation}, issn = {1445-5226}, doi = {https://doi.org/10.1071/IS17052}, url = {https://www.publish.csiro.au/is/IS17052}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak and Backeljau, Thierry} } @article {1781, title = {A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)}, journal = {European Journal of Taxonomy}, volume = {199}, year = {2016}, month = {24/05/2016}, pages = {1-37}, type = {Research Article}, abstract = {

The Thyropygus opinatus subgroup (Diplopoda: Harpagophoridae) of the T. allevatus group in Thailand is revised. Based on a phylogenetic analysis of mtDNA sequence data, it is merged with the T. bifurcus subgroup to form an extended T. opinatus subgroup. Nine new species are described: Thyropygus cimi sp. nov. and T. forceps sp. nov. from Nakhonsrithammarat Province, T. culter sp. nov., T. planispina sp. nov., T. undulatus sp. nov. and T. ursus sp. nov. from Krabi Province, T. mesocristatus sp. nov. from Songkhla Province, T. navychula sp. nov. from Phang-Nga Province and T. sutchariti sp. nov. from Phetchaburi Province.

}, keywords = {DNA barcoding, millipedes, new species, phylogeny, Thailand}, issn = {2118-9773}, doi = {https://doi.org/10.5852/ejt.2016.199}, url = {https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/322}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak and Backeljau, Thierry} } @article {1778, title = {A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae)}, journal = {Zootaxa}, volume = {2980}, year = {2011}, month = {29/07/2011}, pages = {37-48}, type = {Article}, abstract = {

The Thyropygus cuisinieri subgroup of the T. allevatus group is revised. A new species is described from Thailand: T. jarukchusri n. sp. The following species are re-described: T. cuisinieri Carl, 1917, T. carli Attems, 1938, and T. foliaceus (Demange, 1961), new status (elevated from subspecies status under T. cuisinieri).

}, keywords = {millipede, new species, Taxonomy, Thailand}, issn = {1175-5334, 1175-5326}, doi = {https://doi.org/10.11646/zootaxa.2980.1.3}, url = {https://www.mapress.com/zt/article/view/zootaxa.2980.1.3}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak} } @article {1777, title = {A revision of the Thyropygus allevatus group. Part 3: the T. induratus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)}, journal = {Zootaxa}, volume = {2941}, year = {2011}, month = {06/07/2011}, pages = {47-68}, type = {Article}, abstract = {

The Thyropygus induratus subgroup of the T. allevatus group is revised. Three new species are described from Thailand: T. dormiens n. sp., T. laterolobatus n. sp. and T. macrosiamensis n. sp. The following species are redescribed and re-elevated from subspecies status under T. allevatus (Karsch, 1881): Thyropygus induratus Attems, 1936 (= T. punctatus Attems, 1938, new synonym), Thyropygus quietus Attems, 1938, Thyropygus resimus Attems, 1938, and Thyropygus uncinatus (Demange, 1961). Thyropygus siamensis Verhoeff, 1938 (= Thyropisthus ligulus Demange, 1961, new synonym), is re-described. A case of species with identical gonopods but very different body size (T. induratus and T. quietus) is discussed.

}, keywords = {millipede, new species, size difference, Taxonomy, Thailand}, issn = {1175-5334, 1175-5326}, doi = {https://doi.org/10.11646/zootaxa.2941.1.3}, url = {https://www.mapress.com/zt/article/view/zootaxa.2941.1.3}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak} } @article {1775, title = {A revision of the Thyropygus allevatus group. Part 2: the T. bifurcus subgroup (Diplopoda, Spirostreptida, Harpagophoridae)}, journal = {Zootaxa}, volume = {2165}, year = {2009}, month = {22/07/2009}, pages = {1-15}, type = {Article}, abstract = {

The Thyropygus bifurcus subgroup of the T. allevatus group is revised. Four new species are described from Thailand: T. demangei n. sp., T. quadricuspis n. sp., T. richardhoffmani n. sp., from Trang province and T. casjeekeli n. sp., from Krabi province. The other species of the T. bifurcus subgroup, viz., T. bifurcus (Demange, 1986), and T. enghoffi (Demange, 1989), are redescribed.

}, keywords = {millipede, new species, Taxonomy, Thailand}, issn = {1175-5334, 1175-5326}, doi = {https://doi.org/10.11646/zootaxa.2165.1.1}, url = {https://www.mapress.com/zt/article/view/zootaxa.2165.1.1}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak} } @article {1774, title = {A revision of the Thyropygus allevatus group. Part 1: the T. opinatus subgroup (Diplopoda: Spirostreptida: Harpagophoridae)}, journal = {Zootaxa}, volume = {2016}, year = {2009}, month = {20/02/2009}, pages = {17-50}, type = {Article}, abstract = {

The Thyropygus opinatus subgroup of the T. allevatus group is revised. The T. opinatus subgroup corresponds to the genus Cornugonus Demange, 1961, which is formally synonymized under Thyropygus. Eight new species are described from Thailand: T. bearti n. sp., T. brachyacanthus n. sp., T. loxia n. sp., from Suratthani province, T. bispinus n. sp., from Uthaithani and Phrae provinces, T. bispinispatula n. sp., from Chumphon province, T. chelatus n. sp., from Nakhonsrithammarat province, T. cristagalli n. sp., from Phang Nga province and T. erectus n. sp., from Satun province. The other species of the T. opinatus subgroup, viz., T. opinatus (Karsch, 1881), n. comb., T. floweri (Demange, 1961), n. comb., T. implicatus (Demange, 1961), n. comb., and T. inflexus (Demange, 1989), n. comb., are redescribed. New records are given for T. opinatus, T. implicatus and T. floweri. The type locality of T. floweri is shown to be in Thailand rather than Malaysia.

}, keywords = {millipede, new species, Taxonomy, Thailand}, issn = {1175-5334, 1175-5326}, doi = {https://doi.org/10.11646/zootaxa.2016.1.2}, url = {https://www.mapress.com/zt/article/view/zootaxa.2016.1.2}, author = {Pimvichai, Piyatida and Enghoff, Henrik and Panha, Somsak} } @article {1082, title = {A new giant species of the millipede genus Prionopetalum Attems, 1909 from Mt. Kilimanjaro, Tanzania (Diplopoda, Spirostreptida, Odontopygidae)}, journal = {Zootaxa}, volume = {4950}, year = {2021}, month = {31/03/2021}, pages = {561-570}, abstract = {

A new large species of the millipede genus Prionopetalum Attems, 1909, P. nessiae sp. nov. from Mt. Kilimanjaro, Tanzania, is described, with SEM imaging and illustrations of relevant taxonomic characters, and mapped. Comments on the significance of sternum 9 in descriptions of Odontopygidae, and on the intrageneric differences in Prionopetalum, are included.

}, keywords = {new character, new species, Taxonomy}, issn = {1175-5326}, doi = {10.11646/zootaxa.4950.3.8}, url = {https://www.biotaxa.org/Zootaxa/article/view/zootaxa.4950.3.8}, author = {Rosenmejer, Trine and Enghoff, Henrik} } @article {893, title = {A mountain of millipedes IX: Species of the family Gomphodesmidae from the Udzungwa Mountains, Tanzania (Diplopoda, Polydesmida)}, journal = {European Journal of Taxonomy}, year = {2020}, month = {29/06/2020}, pages = {1-35}, abstract = {

A new genus and six new species of the family Gomphodesmidae from the Udzungwa Mts are described, including Pogoro alopias Rosenmejer \& Enghoff sp. nov., Pogoro siren Rosenmejer \& Enghoff sp. nov., Pogoropsis prolixopes Rosenmejer \& Enghoff gen. et sp. nov., Emphysemastix frampt Olsen \& Enghoff sp. nov., Agrophogonus hamulus Olsen \& Enghoff sp. nov., and Agrophogonus pusillokiellandi Olsen \& Enghoff sp. nov. Emphysemastix dracarys Olsen \& Enghoff sp. nov. from Iringa city is described. Descriptive notes are given for Pogoro scharffi Hoffman, 2005, and Agrophogonus mwanihana Hoffmann, 2005. General gomphodesmid gonopod morphology is described and illustrated. A key to Udzungwa gomphodesmids is presented, as well as revised keys to all species of Emphysemastix and Agrophogonus. All gomphodesmid species from the Udzungwa Mts are mapped.

}, keywords = {Eastern Arc Mountains, key, new species, Taxonomy}, issn = {2118-9773}, doi = {10.5852/ejt.2020.675}, url = {https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1005}, author = {Olsen, Sissel Anna and Rosenmejer, Trine and Enghoff, Henrik} } @article {892, title = {A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited {\textendash} a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts and introduction of the first pair of male legs as a source of taxonomic characters}, journal = {European Journal of Taxonomy}, year = {2020}, month = {14/04/2020}, pages = {1-32}, abstract = {

Full title: A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited \– a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae)

A large, newly collected material of the genus Aquattuor from the Udzungwa Mts, Tanzania, has been studied. Two new species are described, viz., A. mollilobus sp. nov. from the Udzungwa Mts and A. nguruensis sp. nov. from the Nguru Mts, Tanzania. Aquattuor claudiahempae Enghoff \& Frederiksen, 2015, is recorded as new for the Udzungwa fauna. Characters from the first pair of male legs are illustrated for all Aquattuor species and are shown to be useful for species discrimination. A new gonopodal difference between A. submajor Enghoff, 2015 and A. udzungwensis Enghoff, 2015 is described and illustrated. Intrageneric relationships and distribution patterns are discussed. A distribution map and a key to all Aquattuor species are presented.

}, keywords = {Eastern Arc, identification key, new species, Taxonomy}, issn = {2118-9773}, doi = {10.5852/ejt.2020.626}, url = {https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/901}, author = {Enghoff, Henrik} } @article {891, title = {A strikingly coloured new giant millipede from Vietnam has copycat in Borneo (Diplopoda, Spirostreptida, Harpagophoridae)}, journal = {European Journal of Taxonomy}, volume = {722}, year = {2020}, month = {20/10/2020}, pages = {1-15}, abstract = {

A new giant species of the subfamily Rhynchoproctinae with strikingly bi-coloured, red-yellow legs contrasting with a black body is described: Alienostreptus bicoloripes sp. nov. from Vietnam. The new species is assigned to the genus Alienostreptus Pimvichai, Enghoff \& Panha, 2010 hitherto comprising one species, A. alienus (Attems, 1936), and differing from other rhynchoproctine genera by having the femoral spine duplicated. Species of this genus share three synapomorphic characters of the subfamily, viz (1) anterior coxal fold forms deep concavity in posterior view, (2) posterior coxal fold very low, and (3) posterior coxal fold with mesal flap. A superficially very similar colourful species from Borneo, also with bi-coloured legs but clearly not belonging to Alienostreptus due to the position of ventral soft pads on male legs, is documented based on photographs.

}, keywords = {new species, Southeast Asia, Taxonomy}, issn = {2118-9773}, doi = {10.5852/ejt.2020.722.1129}, url = {https://europeanjournaloftaxonomy.eu/index.php/ejt/article/view/1129}, author = {Pimvichai, Piyatida and Akkari, Nesrine and Semenyuk, Irina and Constant, Jer{\^o}me and Enghoff, Henrik} } @article {71, title = {A new genus and species of Haplobainosomatidae (Diplopoda: Chordeumatida) from the MSS of the Sierra de Guadarrama National Park, central Spain}, journal = {Zootaxa}, volume = {4347}, year = {2017}, month = {Feb-11-2018}, pages = {492-510}, abstract = {

The chordeumatidan fauna of the Iberian Peninsula is far from being well known, but recent efforts are improving that knowledge. Samplings carried out in the Milieu Souterrain Superficiel (also known as the Mesovoid Shallow Substratum) on several screes of the Sierra de Guadarrama National Park (central Spain) have led to the discovery of a new species of chordeumatidan millipede. This new species belongs to the family Haplobainosomatidae, which is mainly known from the northern Iberian Peninsula, and thus this is the first record of the family in central Spain. However, the extremely simple gonopods of this species are so different from all other known species of the family that it must be placed in a new genus. In this work, a complete description of the species, named as Guadarramasoma ramosae gen. \& sp. nov., with a detailed iconography based on scanning electron microscopy images is provided together with a distribution map and a brief discussion of the implications of this new finding.

}, keywords = {millipedes, new species, orobiome, Sierra de Guadarrama, Subterranean environments}, issn = {1175-5326}, doi = {10.11646/zootaxa.4347.310.11646/zootaxa.4347.3.4}, url = {https://biotaxa.org/Zootaxa/article/view/zootaxa.4347.3.4}, author = {Jos{\'e} D. Gilgado and Ledesma, Enrique and Enghoff, Henrik and Mauri{\`e}s, Jean-Paul and Ortu{\~n}o, Vicente M.} } @article {69, title = {Out of the dark void: Ommatoiulus longicornis n. sp., a new julid from Spain (Diplopoda, Julida) with notes on some troglobiomorphic traits in millipedes}, journal = {Zootaxa}, volume = {4420}, year = {2018}, month = {May-05-2019}, pages = {415-429}, abstract = {

A new millipede species, Ommatoiulus longicornis n. sp. is described from the subterranean habitat in eastern Spain, and the elongation of its antennae is discussed in relation to the habitat where it dwells. We take the occasion to discuss the elongation of appendages as a troglobiomorphic character in millipedes and shed the light on the various, sometimes controversial, hypotheses that could have led to these modifications in millipedes and other subterranean arthropods.

}, keywords = {new species, Schizophyllini, Spain, Taxonomy, troglomorphism}, issn = {1175-5326}, doi = {10.11646/zootaxa.4420.310.11646/zootaxa.4420.3.7}, url = {https://biotaxa.org/Zootaxa/issue/view/zootaxa.4420.3https://biotaxa.org/Zootaxa/article/view/zootaxa.4420.3.7https://biotaxa.org/Zootaxa/article/viewFile/38811/33160}, author = {Akkari, Nesrine and Jos{\'e} D. Gilgado and Ortu{\~n}o, Vicente M. and Enghoff, Henrik} }